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SMITHSONIAN. INSTITUTION UNITED STATES NATIONAL MUSEUM
PROCEEDINGS
OF THE
“NITED STATES NATIONAL MUSEUM
VOLUME 74
SINGS
UNITED STATES GOVERNMENT PRINTING OFFICE WASHINGTON : 1929
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ADVERTISEMENT
The scientific publications of the National Museum include two series, known, respectively, as Proceedings and Bulletin.
The Proceedings, begun in 1878, is intended primarily as a medium for the publication of original papers, based on the collections of the National Museum, that set forth newly acquired facts in biology, anthropology, and geology, with descriptions of new forms and revisions of limited groups. Copies of each paper, in pamphlet form, are distributed as published to libraries and scientific organiza- tions and to specialists and others interested in the different subjects.
The dates at which these separate papers are published are recorded in the table of contents of each of the volumes.
The present volume is the seventy-fourth of this series.
The Bulletin, the first of which was issued in 1875, consists of a series of separate publications comprising monographs of large zoological groups and other general systematic treaties (occasionally in several volumes), faunal works, reports of expeditions, catalogues of type-specimens, special collections, and other material of similar nature. The majority of the volumes are octavo in size, but a quarto size has been adopted in a few instances in which large plates were regarded as indispensable. In the Bulletin series appear volumes under the heading Contributions from the United States National Herbarium, in octavo form, published by the National Museum since 1902, which contain papers relating to the botanical collections of the Museum.
ALEXANDER WETMORE, Assistant Secretary, Smithsonian Institution.
Wasuineton, D. C., November 15, 1929.
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TABLE OF CONTENTS
Aupricu, J. M. Five new parasitic flies reared from beetles in China and India. No. 2753, pp.1-7. October 19, 19281_-_
New genus: Szgelotrozxis. New species: Campylocera hirsuta, Adapsilia flaviseta, Sigelo- troxis parvus, Pexomyia genalis, Centeter unicolor.
Further studies of types of American muscoid flies in the collection of the Vienna Natural History Museum. INO 2764. pp: 1-34.” “Nebrubeyi2o, LO2OM Wake Coe
New species: Copecrypta orbitalis.
New diptera or two-winged flies from South America. NOX 2746. spp l-255) 5 December MrlO28 Dee tp wake n= ON
New genera: Callesthes, Daetaleus, Reedia, Piriona.
New species: Odontomyia tremolerana, Ocnaea gigas, O. falcifer, Ophyra villosa, Hydrotaea fuscisquama Opsidia intonsa, Pachy- ophthalmus ornaticauda, Brachycoma_ ruficauda, Callesthes histrio, Daetaleus purpureus, Incamyia spinicosta, I. striata, I. chilensis, Reedia robusta, Echinopyrrhosia trophocyon, Chaeto- craniopsis argenticeps, Selenomyia virens, S. plena, S. facialis, Piriona fasciculata.
Three new species of two-winged flies of the family Bombyliidae from India. No. 2747, pp. 1-3. October 26,
New species: A phoebantus clauseni, A. seratus, Exoprosopa sipho.
Cusuman, R. A. A revision of the North American Ichneu- mon-flies of the genus Mesostenus and related genera. No. 2761 App. ue58 4, se bruary AOsLO29 Pogo esi. a iuaiee Oogt eye a)
New genera: Agonocryptus, Messatoporus, Acerastes.
New species: Messatoporus rufiventris, M. compressicornis, M. major, Listrognathus multicolor, L. agnatus, Crypturopsis abdominalis, Diapetimorpha rufigaster, D. confederata, D. alabama, D. acadia, Mesostenus melanurus, Polycyrtidea limi- tis, Polistiphaga zonata.
New varieties: Listrognathus albomaculatus rufitibialis, L. a. multimaculatus.
New combinations: Agonocryptus discoidaloides, Messatoporus discoidalis, Mallochia strigosa, Crypturopsis audaz, C. saundersi, C. armatus, Diapetimorpha orba, D. introita, Derocentrus macilentus (Mesostenus) Polycyrtidea pusillus, (Agrypon) Polycyrtidea flavopictus, Acerastes pertinaz.
New name: Diapetimorpha cinctiventris.
Article
8
19
16
1 Date of publication.
VI TABLE OF CONTENTS
Dunn, Emmett Rerp. A new salamander from southern California. No. 2770, pp. 1-3. January 31, 1929?_______
New species: Ensatina klauberi.
Two new frogs from Jamaica. No. 2767, pp. 1-2. December 18, 1928!
New species: Eleutherodactylus orcuttt, E. cunctator.
Se ee
Dyar, Harrison G. Notes and new species of American moths of the genus Scoparia Haworth. No. 2769, pp. 1-9. January 31, 1929!
New species: Scoparia denigata, S. albertalis, S. echo, S. leuco- phthalma, S. phycitinalis, S. molliculella, S. subtersa, S. multi-
facies, S. inexoptata, S. ycarda, S. crassiuscula, S. imparilis, S. vinasalis, S. excursalis, S. ulmaya.
Fosuac, WitutiamM F. Mineralogy and geology of Cerro Mer- cado, Durango, Mexico. No. 2768, pp. 1-27. February D0 1G20 =e aR Ns 1 ay i eR a
GitmorE, CHartes W. Fossil footprints from the Fort Union (Paleocene) of Montana. No. 2750, pp. 1-4. October 6,
New species: Ammobatrachus montanensis.
Kirk, Epwrn. Bathmopterus, a new fossil gasteropod genus from the Silurian of Alaska. No. 2763, pp. 1-4, December ba, GIR te dE a ha ape oh eee ee
New genus: Bathmopterus. New species: Bathmopterus liratus.
The fossil crinoid genus Vasocrinus Lyon. No. 2760, pp. f-10. dentary 20.) LOU * ee re ee ee eter
New genus: Pellecrinus. New species: Vasocrinus turbinatus.
Mattocu, J. R. Notes on some oriental Sapromyzid flies (Diptera), with particular reference to the Philippine species. No. 2751: pp. 1-07” April 22 ao es ae
New genera: Acelyphus, Idiocelyphus, Pleurigona, Euproso- pomyia, Maquilingia, Kerteszomyia, Griphominettia.
New species: Acelyphus politus, A. repletus, Idiocelyphus bakeri, Paracelyphus tesiaceus, Celyphus puncticeps, C. bisetosus, Spaniocelyphus nigrocoeruleus, S. sumatranus, Pleurigona curvinervis, Euprosopomyia maculosa, Pachycerina flaviventris, Phobeticomyia preapicalis, Trypaneoides major, Minettia philip- pinensis, Sapromyza (Sapromyza) maquilingensis, S. (S.) in- versa, S. omei, Maquilingia hirticeps, M. facialis, Trigono- metopsis punctipennis, Kerteszomyia maculifrons, Homoneura (Chaetohomoneura) umbrosa, H. (Neohomoneura) albicosta, H. (N.) setiventris, H. (N.) yerburyi, H. (N.) indica, H. (N.)
Article
25
22
24
23
18
15
1 Date of publication.
TABLE OF CONTENTS
macgregort, H. (Huhomoneura) nigriflua, H. (Homoneura) curta, H. (H.) philippinensis, H. (2.) intermedia, H. (H.) cos- talis, H. (H.) horni, H. (H.) crinita, H. (H.) hirtitibia, H. (H.) dentifera, H. (H.) maquilingensis, H. (H.) opacithorax, H. (#.) piceoides, H. (H.) nigrita, H. (H.) luzonensis, H. (H.) robusta, H. (H.) demeweriu, H. (H.) bakeri, H. (H.) atriceps, H. (H.) neosignata, H. (H.) immaculipennis, H. (H.) fuscobrunnea, H. (#H.) affinis, H. (H.) grahami, H. (H.) sublucida, Gripho- minettia sumatrana.
New subgenera: Minettiella, Xenosapromyza, Luzonomyza, Neo- trigonometopus, Griphoneuroider, Poecilomyza, Minettioides.
New names: Homoneura (Homoneura) piliseta, H. (H.) diacros- tichalis.
MANSFIELD, WENDELL C. New fossil mollusks from the Miocene of Virginia and North Carolina, with a brief out- line of the divisions of the Chesapeake group. No. 2759, Doe fe anwary 4 O29 Tie Se en k Woe ne noe
New species: Fusinus (Buccinofusus) propeparilis, Pisania (Cel- atoconus) burnsi, Columbella (Seminella) smithfieldensis, Epi- tonium smithfieldensis, Fossarus (Isapis) urbannaénsis, Thracia (Cyathodonta) dalli, Pandora (Clidiophora) conradi.
New subspecies: Clathodrillia? belloides rushmerensis, Pecten Eboreus urbannaénsis, Crassatellites (Crassatellites) meridion- alis surryensis, C. (C.) m. urbannaénsis, Phacoides (Lucinoma) contractus murfreesboroénsis.
MarsuHatu, Wituiam B. New fossil pearly fresh-water mus- sels from deposits on the Upper Amazon of Peru. No. 2748, DDAl—7-. VOCtobery sl Wy URO2S doe eases ich pepo IE pep orug aoe
New genera: Prodiplodon, Eodiplodon. New species: Prodiplodon singewaldi, P. bassleri, P. paucarpa- tensis, Hodiplodon gardnerae, E. pebasensis. New fresh-water and marine bivalve shells from Brazil and Uruguay. No.2762,pp.1-7. October 26, 1928 '-
New species: Diplodon jacksoni, D. pilsbryi, Anodontites elfa, A. mansfieldi, M ycetopoda felipponei, Corbula uruguayensis, Nucula felipponei, N. uruguayensis.
Miuurr, Gerrit S., Jr. The gums of the porpoise Phoccenoi- des dalli (True). No. 2771, pp. 1-4. January 26, 19291__- Price, EmmMerr W. Two new nematode worms from rodents. Nos 2 749. ppt l-Syie November 10, NO28 Mauka NT 2 8
New species: Syphacia thompsoni, Heligmostrongylus hassalli.
Scuwartz, Bensamin. A new species of trichostrongylid worm of the genus Cooperia from the carabao in the Philip-
pine Islands, with a review of the genus. No. 2765, pp. 1-5. ING errno ety 2G SY os a apie cape ati 1 edo let ee A ey 3 ed
New species: Cooperia nodulosa.
1 Date of publication.
VII
Article
14
17
VIII TABLE OF CONTENTS
SHANNON, Eart V. Miargyrite silver ore from the Rands- burg district, California. No. 2766, pp. 1-10, January 31,
Tschermigite, ammoniojarosite, epsomite, celestite, and paligorskite from southern Utah. No. 2758, pp. 1-12. JANE Aly Wee peice he ) eee ed Ge eee
Taytor, E. Leonarp. Moniezia, a genus of cestode worms, and the proposed reduction of its species to three. No. 2754, pp t-0 Ti Nosembper 24, 1928 oe ee
Topp, W. E. Crypr. A revision of the wood-warbler genus, Basileuterus and its allies. No. 2752, pp. 1-95. April 26,
New genus: Phaeothlypis.
New subspecies: Phaeothlypis semicervina annexa, Basileuterus castaneiceps chapmani, B. coronatus notius, B. c. elatus, B. c. regulus, B. signatus flavovirens.
TREADWELL, AARon L. A new polychaetous annelid of the genus Phyllodoce from the west coast of Costa Rica. No. 2157;.DP. lS.) NOVEMDOR.NU, doom ee ee
New species: Phyllodoce nicoyensis.
Van Duzer, M.C. Tropical American Diptera or two-winged flies of the family Dolichopodidae from Central and South America. No. 2755, pp. 1-64. February 9, 1929!_______
New genera: Keirosoma, Oedematopus.
New species: Psilopus bilobus, P. longipes, P. semicomatus, P. planipes, P. mensor, P. bicoloripes, P. longipennis, P. cylindri- cus, P. angustatus, P. semiciliatus, P. panamensis, P. nitidi- cauda, P. flaviannulatus, P. clavatus, P. simulans, Diaphorus curvispina, D. perplerus, D. varipes, Asyndetus flavitibialis, Chrysotus contractus, C. flavimaculata, Argyra violacea, Keiro- soma albicinctum, Sympycnus imperfectus, S. longipes, S. fili- formis, S. bipilus, Nothosympycnus unipilus, N. furcatus, Neu- rigona banksi, N. maculipennis, Thinophilus panamensis, Me- detera bella, M. pollinosa, M. scaura, M. pallidicornis, M. varipes, M. flaviseta, Thrypticus acuticauda, Oedematopus cras- sitibia, Paraclius angustipennis, P. albitalus, P. simplex, P. parvicauda, P. aldrichi, P. latitibia, P. albimanus, P. argen- timanus, P. frontalis, P. terminalis, P. acuticornis, Polymedon transversus, P. nigricornis, P. partitus, P. rubiginosus, P. no- tatus, Sarcionus maculatus, Pelastoneurus maculitibia, P. acuti- cauda, P. fuscipennis, P. peciinatus.
Wricut, A. H. Synopsis and description of North American tadpoles. No. 2756, pp. 1-70. June 4, 1929!_________-_-
1 Date of publication.
Articl.
21
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LIST OF ILLUSTRATIONS
PLATES
NEW FOSSIL PEARLY FRESH-WATER MUSSELS FROM DEPOSITS ON THE UPPER AMAZON OF PERU
By William B. Marshall Facing page
1) (New fresh-water mussels fron) Peru st S20 Ae Yeo 8 Two NEW NEMATODE WORMS FROM RODENTS By Emmett W. Price Lusypnaca thompsonr, new speciess! Sao e ia Wael Be ei ee 6 2. ‘Heligmosirongylus hassall:, new species! s_- . = 225-- 222 225s 22-23. 6 FossIL FOOTPRINTS FROM THE Fort UNION (PALEOCENE) OF MontTaNa By Charles W. Gilmore 1. Type slab of Ammobatrachus montanensis__-.__...._-._..----.-- .-- 4 2-3. Part of type slab of Ammobatrachus montanensis_.__..--.---------- 4 Nores ON SOME ORIENTAL SAPROMYZID FLIES (DIPTERA), WITH PARTICULAR REFERENCE TO THE PHILIPPINE SPECIES By J. R. Malloch i ‘Details of Celyphinae and Sapromyzinae= 2 Pek Aer ae: 89 Pre DeLAtistOL Sa prOmly ZINAek Als we eo ee A eae 90 Se ELAS TORMTOITONEUN nna oa wt ores ee eae ea GY a 91 Aol El ypOpy eid, OL SPeCles Ol -HOmMONeUTa coe ee 92, 93 6. Hypopygia and apical genital segments of species of Homoneura___-_-- 94 MONIEZIA, A GENUS OF CESTODE WORMS, AND THE PROPOSED RE- DUCTION OF ITS SPECIES TO THREE By E. Leonard Taylor ZV aniationstinMontecta ex pansa ss ue ec iae! ee oy a ae 10 3. Variations in Moniezia expansa and M. planissima__.-_-------------- 10 AV aMananssin MOonveZta ex PAUS@ an se aia. be othe Ge a es 10 5. Variations in Moniezia expansa and M. planissima_.._-------------- 10 TRoPIcAL AMERICAN DIPTERA OR TWO-WINGED FLIES OF THE FAMILY Do.icHOPpoDIDAE FROM CENTRAL AND SouTH AMERICA By M. C. Van Duzee 1-2. Details of the flies of the family Dolichopodidae___._______..------ 64
LIST OF ILLUSTRATIONS
SYNOPSIS AND DESCRIPTION OF NorRTH AMERICAN TADPOLES
By A. H. Wright
Facing page 1-5. Mouth parts of North American tadpoles_...-.-..-.------------- 6—9, Lateraliaspects.of livectadpoless=2=~.- = 2-2-2 es ee
NEW FOSSIL MOLLUSKS FROM THE MIOCENE OF VIRGINIA AND NorTH CAROLINA, WITH A BRIEF OUTLINE OF THE DIVISIONS OF THE CHESAPEAKE GROUP
By Wendell C. Mansfield
1-5. New fossil mollusks from the Chesapeake group------------------
THE FOSSIL CRINOID GENUS VAsocRINUS LYON
By Edwin Kirk
1-2. Species of fossil crinoid genus Vasocrinus_-..--------------------
NEw FRESH-WATER AND MARINE BIVALVE SHELLS FROM BRAZIL AND URUGUAY
By William B. Marshall
1—4.. Bivalve shells from: Brazil and Uruguay ~- =. 2--------2-=------
PON
Pwr
BATHMOPTERUS, A NEW FOSSIL GASTEROPOD GENUS FROM THE SILU- RIAN OF ALASKA
By Edwin Kirk A new fossil gasteropod from the Silurian of Alaska_______----------
A NEW SPECIES OF TRICHOSTRONGYLID WORM OF THE GENUS CoopE- RIA FROM THE CARABAO IN THE PHILIPPINE ISLANDS, WDE: “A: REVIEW OF THE GENUS
By Benjamin Schwartz
s Cooperta nodulosa= New Species $2 s= an a eee eee ee ene ot
MINERALOGY AND GEOLOGY OF CERRO Mercapo, DurANGO, MEXICO By William F. Foshag
Cerro Mercado from the south, the bold outcrops and iron ore------- Views of CemrobMercadots {2 <3. ee ee ee ee Breccia partially replaced" by diopside= =." "> "===.- 2-5 5---= sae Breccia partially replaced by calcite and magnetite - ----------------
THE GUMS OF THE PORPOISE PHOC@NOIDES DALLI (TRUE)
By Gerrit 8S. Miller, jr.
Teeth of a Phocaena and b Phocoenoides_-_-_------------- esc ae See Jaws of Phocoenoides <<. wie ea ee eee a ee Guns of Phocoendides, lower jaw 3.- 4-22. 22s ee eee
Gums;ofsPhocoenvides, Upper (aWes. ] tone a eens
70
12
16
SPP Pp
LIST OF ILLUSTRATIONS
TEXT FIGURES NEW DIPTERA OR TWO-WINGED FLIES FROM SouTH AMERICA
By J. M. Aldrich
. Daetaleus purpureus. Genital segments and genitalia of male from left side: 1, 2, genital segments; if, inner forceps; of, outer forceps; (ols SO) TNS4 OY eS re 00 OY 2) GT Aa amp a 2 ae aN BR eS By . Piriona fasciculata, new species. Hind leg of male_________________
FossIL FOOTPRINTS FROM THE Fort UNION (PALEOCENE) OF Montana
By Charles W. Gilmore
. Ammobatrachus montanensis. Diagram of trackway__________-____-
A REVISION OF THE WOOD-WARBLER GENUS BASILEUTERUS AND ITS ALLIES
By W. E. Clyde Todd
. Diagram of the phylogeny of Basileuterus and allied genera__________
A NEW POLYCHAETOUS ANNELID OF THE GENUS PHYLLODOCE FROM THE WEST COAST OF Costa RICA
By Aaron L. Treadwell
1-3. Phyllodoce nicoyensis, new species: 1, prostomium and _ proboscis;
2 pSPATADOCIMIN eoaISe baler Be ANN a aioe Ae Se ee shee
TSCHERMIGITE, AMMONIOJAROSITE, EPSOMITE, CELESTITE, AND PALIGORSKITE FROM SOUTHERN UTAH
By Earl V. Shannon
ieCelestiteserystal. Panguitch Utah «poe o ees hea ee
A REVISION OF THE NortTH AMERICAN ICHNEUMON-FLIES OF THE GENUS MESOSTENUS AND RELATED GENERA
By R. A. Cushman
1. Head of: a, Listrognathus albomaculatus (Cresson), dorsal view; b, same,
lateral view of lower part to show occipital carina (oc) and hyposto- mal carina (he); c, Listrognathus agnatus Cushman, lateral view of lower part; d, Polyaenus spinarius (Brullé), dorsal view; e, Mesos- tenus leucopus Ashmead, dorsal view; f, Polycyrtus neglectus Cush- man, dorsal view; g, Polycyrtidea limitis Cushman, dorsal view _ _-__-
2. First abdominal segment of female: a, Agonocryptus discoidaloides
(Viereck), lateral view; b, Messatoporus discoidalis (Cresson), lateral view; c, Mallochia agenioides Viereck, dorsal view; d, same, lateral view; e, Listrognathus albomaculatus, lateral view; f, Crypturopsis audax (Cresson), dorsal view; g, Mesostenus leucopus Ashmead, dorsaliviews hysame) lateraloviewss.ee 2.22 lees ae Fee
3. Propodeum of female: a, Crypturopsis abdominalis Cushman; b, Agono-
cryptus discoidaloides (Viereck); c, Messatoporus discoidalis (Cres- son); d, Listrognathus albomaculatus (Cresson); e, Listrognathus agna- .tus Cushman; f, Diapetimorpha introita (Cresson); g, Derocentrus
XI
Facing page
XII LIST OF ILLUSTRATIONS
Facing page
macilentus (Cresson); h, Polycyrtus neglectus Cushman; i, Acerastes pertinaxz (Cresson); j, Mesostenus thoracicus (Cresson); k, Polyaenus spinarius (Brullé); l, Mallochia agenioides Viereck; m, Polistiphaga enigig Gushinal. "22. Soe ee ae ee So. oe oe ee
4, Hind trochanter and femur of: a, Derocentrus macilentus (Cresson) ; b,. Mesostenus, thoracicus, Cresson... - - -. nga ee a3 esa - ==
5, Fore wing of Acerastes perlinas. (Cresson) .. . 22-3 a = 6. The areolet of: a, Agonocryptus discoidaloides (Viereck); b, Messa- toporus discoidalis (Cresson); c, Messatoporus major Cushman; d, Mallochia agenioides Viereck; e, Listrognathus albomaculatus (Cres- son); f, Drapetimorpha orba (Say); g, Diapetimorpha introita (Cres- son); h, Crypturopsis teranus (Ashmead); 1, Polyaenus spinarius (Brullé); 7, Mesostenus thoracicus Cresson; k, Polycyrtus neglectus Cushman; 1, Derocentrus macilentus (Cresson); m, Polycyrtidea limitas Cushman; n, Acerastes pertinax (Cresson); 0, Polistiphaga fete: ( Crassan) Leis) ere aes sot ee er _ Se nea &
. Agonocryptus discoidaloides (Viereck)--._------------------------- « Messatoporus discotdatis (Cresson). - 2. ee ee
on
FURTHER STUDIES OF TYPES OF AMERICAN MUSCOID FLIES IN THE COLLECTION OF THE VIENNA Naturaut History Museum
By J. M. Aldrich
i
. Head of Cuphocera macrocera Wiedemann, male__------------------ . Lepidodexia tetraptera Brauer and Bergenstamm: 1, left lateral view of genital segments of male; 2, rear view of forceps; 3, rear view of penis; 4, tip of penis; 5, rear view of accessory plate_-._----------
bo
A NEW SPECIES OF TRICHOSTRONGYLID WORM OF THE GENUS CoopERIA FROM THE CARABAO IN THE PHILIPPINE ISLANDS, WITH A REVIEW OF THE GENUS
By Benjamin Schwartz 1. Cooperta.noedulosa. Top view, of head .=---.+- 2-2-2 - 2-2
MIARGYRITE SILVER ORE FROM THE RANDSBURG DISTRICT, CALIFORNIA
By Earl V. Shannon
—
. Crystal of miargyrite thick tabular on c (001)---------------------- . Crystal of miargyrite thick tabular on a (100) --------------------- . Crystal of miargyrite tapular ‘onc’ (O01) ~2 =) = Se eeee* Seu see ee
Ww bd
MINERALOGY AND GEOLOGY or Cerro MeErcapo, DURANGO, Mexico
By William F, Foshag
Map, or erro TetesdG. os oe. eee ee ee een . Krystal aire O) BAUbes = o-oo te ae a en ores . SOorvatal Rabie OF BUSING. - oo ok eee oe ie Sere
wne
or or
25
32
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11 14
NEW DIPTERA OR TWO-WINGED FLIES FROM SOUTH AMERICA
By J. M. Axpricu
Associate Curator, Division of Insects, United States National Museum
The following paper contains descriptions of 20 new species and 4 new genera of Diptera from South America. The types of all the species, except that of Dactaleus purpureus, are in the United States National Museum.
As the paper was going through the press it became possible to include as paratypes some material collected in Chile and Argentina by Edwards and Shannon. I hope to take up the bulk of their muscoid collections in a later paper.
Family STRATIOMYIIDAE Genus ODONTOMYIA Meigen
Odontomyia MEIGEN, Illiger’s Mag., vol. 2, 1803, p. 265.
OCDONTOMYIA TREMOLERANA, new species
One of the largest species (male 15 mm., female 16 mm.) ; thorax ereen throughout except for three dorsal black stripes, which in the males join together just before the scutellum; the central stripe is greatly widened in front. Between the front and middle coxae, along the median line is an oblong area which is reddish yellow. Abdomen green with a dorsal narrow, sharply defined black median stripe on second and third segments, spreading vaguely out over the fourth and fifth; in the female the abdomen is very broad and there is a distinct trace of a brown stripe on each side halfway between the middle and the margin. Coxae and femora green; tibiae and tarsi reddish.
Head of male green below and behind, rather strongly carinate below the antennae; labella large, black; eyes bare, hardly contiguous, with an area of small facets occupying the lower fourth. First two antennal joints reddish, short, of equal length, third joint broken off.
The female has small eyes, as usual, the front green with a round- ish black spot covering the ocelli, and a black band from eye to eye
No. 2748.—PROcEEDINGS U. S. NATIONAL Museum, VoL. 74, ART. 1 2605—28 1 1
2 PROCEEDINGS OF THE NATIONAL MUSEUM VOL, 74
passing just above base of antennae, the color deepest in the middle and close to eye.
Wings hyaline, the costal margin yellow; third vein forked at tip; three veins from discal cell which curve forward and become evanescent.
Described from two males taken at Menafra, Rio Negro, Uru- guay, December 15, 1914; and one female from Montevideo; all re- ceived from Juan Tremoleras, to whom the male paratype has been returned.
Type.—Female, Cat. No. 29367, U.S.N.M.
The nearest related described species appear to be Odontomyia quadrilineata Macquart* and O. histrio Walker. Both are large species like this; the former, however, has a black band across the middle of the front in the female, and the latter has a spot on each side marking the beginning of the band; in both the description of the abdomen fails to indicate such a narrow black median stripe as in the present species.
Family CYRTIDAE Genus OCNAEA Erichson
Ocnaea EricHson, Entomographien, 1840, p. 155.—CoLr, Trans. Amer. Ent. Soc., vol. 45, 1919, p. 23, definition and key to North American species. The genus includes 13 known species, all American and mostly tropical; four, however, occur in the southwestern part of the United States, namely, coerulea, helluo, and loewi in Texas, and flavipilosa in Arizona. The two species here described are from Ecuador, and among the largest of the genus, the specimens measuring from 12 to 17 mm. in length. Only calida and grossa approach this size, and the latter, having bare eyes, is probably better placed in Apelleia Bellardi.
OCNAEA GIGAS, new species
Male.—Chestnut brown, second abdominal segment lighter, third to sixth segments above except at sides much darker in two of the three specimens, blackish with slight bluish tinge; venter with the broad sternites variably infuscated; thoracic dorsum subshining, with or without a pair of lighter submedian lines diverging forward; femora and tibiae black, the tarsi brown. Antennae blackish, in- serted just in front of the large posterior ocelli (the front ocellus absent), basal joints large, third long, sickle shaped, slender and curved downward, reaching below the head, without hairs on any part. It is twice as long as in fwmens, figured by Walker. The
1 Hist. Nat. Dipt., vol. I, 1834, p. 248, locality South America. ?Dipt. Saund., 1851, p. 77, locality Brazil.
ART. 1 NEW DIPTERA FROM SOUTH AMERICA——-ALDRICH 3
compound eyes literally cover the whole head except the flattened hind surface and a triangle above where the ocelli and antennae are situated. The two posterior ocelli are large and prominent, a trans- verse groove just anterior to them. Pile of eyes dense, dark in front, more pale yellow at sides. Proboscis visible but minute. Mesonotum covered with dense glistening pale yellow pile, the ground color mostly dark yellow with or without faint lighter stripes. Hind calypters translucent brownish yellow with well- defined brown rim and yellow fringe. Pile of abdomen yellow, much less conspicuous than on mesonotum. Wing small, tinged with yellow; the second and third veins fork just before the anterior crossvein, which is at the extreme base of the discal cell; an extra crossvein in the first posterior cell just beyond tip of discal, the second section of the first posterior closed and petiolate; all of the veins reach the margin; fourth posterior closed and with compara- tively long petiole which is sometimes equal to the vein forming the apical boundary of the cell.
Length, 13 to 17 mm.
Described from three males collected at Mera, Ecuador, in February, 1923, by F. X. Williams; they were received from the Hawaiian Sugar Planters’ Association, to which one paratype is returned. One specimen has notably paler thorax and abdomen than the others, having no distinct darker abdominal marking.
Type.—Male, Cat. No. 40987, U.S.N.M.
OCNAEA FALCIFER, new species
Male.—Head, thorax, legs, and abdomen black or brown, the second abdominal segment yellow on apical half or more, sides of second, third, and fourth, and posterior bands on the sternites also yellow; third antennal joint brownish yellow, long, slender, and sickle shaped as in gigas, but lighter in color; eyes with dense, long black pile; thorax with dense blackish pile, which has a reddish tinge when viewed from in front. Hind calypter deep brown, with black border and brown fringe. Legs black, the tarsi a little lighter. Wings small, with yellow tinge; venation as in gigas, but the second section of the first posterior cell is wide open in one specimen, narrowly closed in the other, and the fourth posterior cell is closed near margin in three of the four wings, wide open in the other. This open fourth cell is on the other fly from the open first.
Length, 12 to 14.7 mm.
Described from two males, collected with the specimens of gigas. The paratype is returned to the Hawaiian Sugar Planters’ Asso- ciation.
Type.—Male, Cat. No. 40988, U.S.N.M.
4 PROCEEDINGS OF THE NATIONAL MUSEUM VOL, 74
This species is so much like gigas in nearly all structural details that I have considered the possibility that one is a color form of the other. It is apparent that the color and venation are variable, but the differences seem to be suflicient to make the species valid.
Family ANTHOMYIIDAE Genus OPHYRA Robineau-Desvoidy
Ophyra Ropinrau-Desvorpy, Myodaires, 1830, p. 516—RonpDANI, Dipt. Ital. Prod., vol. 6, 1877, p. 34—VAN DER WULP, Biologia, Dipt., vol. 2, 1896, p. 323.—Srermn, Archiv Naturgesch., vol. 83, 1917 (1919), p. 180.—Srevy, Anthomyides de France, 1923, p. 277.—MaAttocH, Ann. Mag. Nat. Hist., vol. 11, 1923, p. 664.
Before describing a new species from Chile, I will notice the species already known from South America. All the extra-European
species are listed by Stein in the article just cited by him.
OPHYRA AENESCENS Wiedemann
Anthomyia aenescens WirDEMANN, Auss. Zweifl., vol. 2, 1830, p. 435. Ophyra argentina Bieot, Annales, 1885, p. 302.—Gie1io-Tos, Ditt, del Mess., vol. 4, 1895, p. 26—VAN DER WULP, Biologia, Dipt., vol. 2, 1896, p. 323. Ophyra aenescens, STEIN, Berlin Ent. Zeitschr., vol. 42, 1897, p. 170; Ann. Mus. Nat. Hung., vol. 2, 1904, p. 451, and vol. 16, 1918, p. 234; Arch. Naturgesch. vol. 76, 1911, p. 100, and vol. 84, 1918 (1920), p. 42; Zeitschr. Hym. Dipt., vol. 7, 1907, p. 212—BisHopp and LAAKE, Journ. Agric. Research, vol. 21, 1921, p. 729.
Ophyra carbonaria SHANNON and Det Ponte, Rev. Inst. Bacteriologico, Buenos Aires, vol. 4, No. 5, 1926 (1927), pp. 20, 30.
This species may be distinguished from the others by having yellow palpi and black knobs on the halteres; it has a bronze tinge. It has been reported from Northern Chile by Stein, 1911 (Caldera, Anto- fagasta, Arica), and is known from Argentina, Brazil, Peru, etc., north to the southern part of the United States. O. carbonaria was recently described from Argentina. Bishopp and Laake, 1921, report that in their experiments they found the adult to fly more than four miles at the maximum from a point of liberation. I have received no Chilean specimens of this species.
OPHYRA SETIA Walker
Anthomyia setia WaLkKER, List Dipt. Ins., vol. 4, 1849, p. 956. Ophyra setia Stern, Zeitschr. Hym. Dipt., vol. 1, 1901, p. 208.
This was described from a single female, collected by Darwin in the Galapagos Islands. Stein on examining the type in the British Museum reported that it is very like O. aenescens except in having yellowish halteres. No other specimens have been reported, and the male is unknown. However, the United States National Museum has four males from the Galapagos Islands of undoubted O. aenescens,
ART, 1 NEW DIPTERA FROM SOUTH AMERICA—-ALDRICH 5
the knobs of the halteres being brown. It may be doubted whether O. setia is specifically distinct from O. aenescens.
OPHYRA VIRESCENS Macquart
Ophyra virescens Macquart, Dipt. Exot., vol. 2, part 3, 1848, p. 321 (sep. p. 164).
Briefly described from a female taken at Guaratuba, Brazil. The type is probably no longer in existence, and the identity of the species is very uncertain. ‘The characters mentioned are: shining green; face with whitish pollen; antennae and legs black; calypters pale yellow; length 6 millimeters. Macquart also says, “thorax a bandes noirs,” which as I understand his terms may mean either black longitudinal stripes or transverse bands on the thorax. If this character is well developed, in either sense, it ought to make the species recognizable. So far it remains unrecognized.
OPHYRA CUTILIA Walker
Anthomyia cutilia WALKER, List. Dipt. Ins., vol. 4, 1849, p. 954. Ophyra cutilia StTe1n, Zeitschr. Hym. Dipt., vol. 1, 1901, p. 194.
Described from Montevideo, only the female being known. Stein has added a few characters from his examination of the type. The most prominent feature is the blackish calypters. I have a male which I identify as this species, as the calypters are the same, and the deep blue color of the female occurs in this male overlaid with such pruinescence as usually distinguishes the sex in this genus.
Color dark blue; the head, pleurae, legs, antennae, and palpi are black; calypters brown, the hind one with blackish rim and long brown fringe. Wings uniformly infuscated.
Male.—Front very narrow, not much wider than front ocellus, still the linear shining parafrontals are separated by a slender median stripe. Lunule silvery, lower parafrontal shining black, quite narrow; the parafacial with brown pollen beginning at about the tip of the second antennal joint. Cheek about one-seventh of the eyeheight. Frontal bristles beginning below the middle of the space between ocelli and lunule; antenne dark brown, the arista distinctly pubescent on basal third, slightly thickened at base, hind edge of eye scarcely at all emarginate in side view. Hairs of back of head all black.
Thorax almost black, subopaque; acrostichals 1,1 (anterior small) ; dorsocentrals 2, 4; intraalar 2, supraalar 1 (no prealar) ; presutural 1; sternopleura with a small bristle anteriorly and one large one behind, acrostichal hairs in about four rows. Hind calypter long and with unusually heavy fringe. Halteres yellow with blackish knob.
6 PROCEEDINGS OF THE NATIONAL MUSEUM VoL. 74
Abdomen with rather dense and erect hair, rather flat toward the base with thin brownish pollen when viewed from behind on which is the beginning of a median dark line. Front tibia without outer flexor bristle, middle tibia with two bristles behind, the upper at the middle, hind tibia with a very distinct calcar below the middle, and on the outer flexor side two bristles below the middle. Middle femur with about a dozen truncated bristles on the under side near base, on the hind side with a dense slanting row of small bristles extending the whole length. Hind femur on the outer side with a distinct row of bristles the whole length, becoming more dorsal toward the tip, also two or three stubby truncate bristles near base on under side and a partial row of bristles on the outer flexor side beyond middle. Claws and pulvilli short. Wings rather narrow. evenly infuscated.
Length, 6.4 mm.
Described from one male, Sao Paulo, Brazil, Dr. A. Lutz.
OPHYRA VILLOSA, new species
Shining blue green with black legs, antennae and palpi. Wings hyaline. Calypters white. Hind tibia with long villosity.
Male.——¥ront nearly as wide as ocellar triangle, the shining black distinctly punctured parafrontals separated by the median stripe which at narrowest is as wide as one parafrontal. The frontal bristles begin just above the middle of the space between the ocelli and lunule, the latter silvery, very conspicuous. Parafacials nar- row with dull gray pollen which extends across the face. Cheek about one-tenth the eyeheight. Hind edge of eye distinctly emargi- nate along the middle. Mesonotum with rather dense erect hairs, entirely shining. Before the suture there are about four rows of delicate rather long acrostichal hairs, but the remaining surface is so hairy that they are difficult to distinguish. Dorsocentral bristles more or less hair-like except the hindmost pair. Anterior sterno- pleural very minute or absent, the calypters white with white rim and fringe, the hind one projecting considerably but not so Jong as in O. cutilia. Halteres black the stem yellow toward base.
Abdomen wholly shining with long erect pile except in the middle of the dorsum. Legs black, claws and pluvilli not elongated; front tibia without a bristle on the outer flexor side. Middle tibia with two bristles on hind side. Hind tibia with long, erect, but not dense villous hairs or bristles in two rows on the outer and inner flexor sides, the intervening space with few hairs. Wings hyaline, third and fourth veins very slightly convergent.
Length, 4.2 to 4.6 mm.
ART. 1 NEW DIPTERA FROM SOUTH AMERICA—ALDRICH cL
Described from two males, collected at Perales, Chile, February 1925, by Father Anastasio Pirion, received from Alfredo Faz, who has retained some additional specimens.
Type.—Male, Cat. No. 28897, U. S. N. M. Genus HYDROTAEA Robineau-Desvoidy
Hydrotaea RopinEAu-Desvoidy, Myodaires, 1830, p. 509.
HYDROTAEA FUSCISQUAMA, new species
Deep black, including palpi, antennae and legs; closely allied to H. dentipes Fabricius, but having infuscated calypters with black rim and fringe, and the abdomen with light blue pruinosity.
Male.—Front including parafrontals black, about as wide in mid- dle as the ocellar triangle; parafacials silvery from the antennal insertion down, becoming brown on lower part; lunule small, golden. Two or three upturned bristles on the cheek next to the vibrissae.
Thorax rather shining black, only when viewed directly from be- hind is there is trace of brown pruinose stripes anteriorly. Acrosti- chal 3,5; dorsocentral 2,4; humeral 3; posthumeral 1; presutural 1; notopleural 2; supraalar 3 (small except middle one) ; intraalar 3; postalar 2; sternopleural 1,1 (some posterior large hairs almost like bristles). Halteres black. Calypters, as noted above, the posterior projecting far beyond anterior.
Abdomen with striking light blue color on black ground, only a trace of median dark stripe. The blue pruinosity extends on venter to the membrane, and even on the sternal plates.
Legs shining black, pulvilli white and not enlarged. Front femur with the usual two thorns below near tip; on the outer flexor side below near base are five long bristles, on the inner side just before the thorn is a row of about six short depressed spines. Front tibia irregularly flattened and concave on flexor side, on outer flexor side near tip with elongated brush-like series of erect hairs. Middle femur with two rows of bristles on front side of basal half, the lower long; a uniform row of small slanting bristles on hind side, and on lower hind side a row of long straight spines on basal half. Middle tibia with rather dense, short, suberect hairs on whole length of front side, on hind side two bristles near middle. Hind tibia straight, not provided with a keel-like brush of short hairs on flexor side at apex, but with a long calcar below middle and on outer side a fairly regu- lar row of cilia.
Wings distinctly brown, the fourth vein converging toward third precisely as in H. dentipes.
Female.—Front with the usual cruciate bristles. Parafacials col- cred as in the male, but a little wider. Thorax asin male. Abdo-
8 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
men ovate but colored as in male. Legs plain as in female ZH. dentipes. Wings and calypters less infuscated than in male, but the latter with dark rim and fringe.
Length, male 6.3 mm., female the same.
Described from one male and one female, collected at Perales, Quilpue, Chile, by Father Anastasio Pirion, on January 25. Received from Alfredo Faz, who has other specimens not seen by me.
Type.—Male, Cat. No. 28876, U.S.N.M.
Family SARCOPHAGIDAE Genus OPSIDIA Coquillett
Opsidia CoqurmLtETT, Journ. New York Ent. Soe., vol. 3, 1895, p. 102.— ALLEN, Proc. U. S. Nat. Mus., vol. 68, art. 9, 1926, p. 41.
OPSIDIA INTONSA, new species
Male——Front narrowest on the lower part where it measures half the head width, the eyes approximate each other still more across the face. Head covered with dark plumbeous pollen and unusually hairy and bristly; frontal stripe almost the same color as the parafrontal, the latter with a somewhat double row of bristles and two proclinate orbitals; parafacials broad with numerous coarse hairs and a diagonal row of bristles above the transverse impression. Front margin of the head in profile is shorter and more vertical than in O. gonioides, genotype of Opsidia, and the third antennal joint is much shorter. Antennae black, the base somewhat reddish, the third joint less than three times the second; arista rather short, the penultimate joint not elongated; antennal depression quite deep, ending, however, a Iit- tle above the mouth; vibrissae not well developed; palpi brown, of ordinary structure; proboscis slender at base, the tip broken off.
Thorax blackish with a little thin pollen forming more distinct cinereous spots on the humeri, postalar declivity and margin of scutellum. Anterior acrostichal 3, posterior numerous and hairlike. _Pleurae shining black.
Abdomen subshining black with thin whitish pruinosity denser on the bases of the segments, except the first, and forming a larger spot on the middle of each. First, second, and third segments with a single pair of median marginals; fourth segment with two or three pairs.
Legs black, middle tibia with one bristle on the outer front side; hind tibia with about a dozen forming a rather even row on the outer hind side and one large in the middle of the row.
Wings subhyaline, the base yellow, veins elsewhere black; first posterior cell open far before the apex; bend of fourth vein with a distinct fold but no fork; hind cross vein joining the fourth more
ART. 1 NEW DIPTERA FROM SOUTH AMERICA—ALDRICH 9
than two-thirds of the way from the small cross vein to the bend; last section of the fifth vein one-third the preceding. Veins bare except the third, which has four or five hairs at base.
Length, 6.2 mm.
Described from one male collected at Santiago, Chile, by Alfredo Faz.
Type.—Male, Cat. No. 28908, U.S.N.M.
Genus PACHYOPHTHALMUS Brauer and Bergenstamm
Pachyophthalmus BRAUER and BERGENSTAMM, Zweifl. Kais. Mus., pt. 4, 1889, p. 117; pt. 6, 1893, pp. 170, 195.—ALtEN, Proc. U. S. Nat. Mus., vol. 68, art. 9, 1926, p. 7.
PACHYOPHTHALMUS ORNATICAUDA, new species
Male.—Front 0.21 of head width, covered with yellowish pollen and having the usual double row of bristles to the antennae. Parafacial with slightly paler pollen, the facial ridges with a few small bristles above the vibrissae; antennae black, the third joint a little longer than the second; arista short; palpi black, proboscis short.
Thorax shining black with very striking white pollinose stripes, one on each side of the middle, and another from the humerus to the root of the wing having a more yellow tinge behind the humerus. When viewed from behind there is also an incomplete stripe from the suture to the postalar declivity above the root of the wing. The white stripes next to the middle reach to the apex of the scutellum. Pleura very pollinose, a black stripe from the humerus passing below the root of the wing and another indefinite stripe along the upper edge of the sternopleura.
Abdomen black with a paler pollinose stripe on each side of the median line partially divided into spots and composed of pollen which shows some brown reflections; the first three segments are broadly gray pollinose below and on the sides, that on the sides forming a silvery spot on the second and third segments, the fourth segment deep golden pollinose, especially well colored on the sides, the hind edge wider in the middle, shining black. Genital segments black; inner forceps shining black, the outer yellow.
Legs black, the claws and pulvilli elongate. Wing hyaline with the usual venation for the genus.
Length, 5.2 mm.
Described from one male, collected at Perales, Chile, by Father Pirion and transmitted to the U. S. National Museum by Alfredo Faz.
Type—Male, Cat. No. 28909, U.S.N.M.
2605—28 2
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The species has some resemblance to P. trivittata Townsend, de- scribed from Peru,* but the general color of the thorax and abdomen is much blacker and the pollen is rather bluish instead of yellow so that the deep golden or almost orange pollen of the fourth seg- ment makes a very strong contrast.
Genus BRACHYCOMA Rondani
Brachycoma Ronpani, Dipt. Ital. Prod., vol. 1, 1856, p. 69.
Laccoprosopa TOWNSEND, Trans. Amer. Ent. Soc., vol. 18, 1891, p. 365; Ins. Ins. Menst., vol. 3, 1915, p. 116.
Bombobrachycoma TowNseEnD, Ins. Ins. Menst., vol. 6, 1918, p. 157. -
BRACHYCOMA RUFICAUDA, new species
Entirely black except the fifth abdominal segment in female and second genital segment of male, which are bright reddish-yellow.
Male.—F ront at narrowest one-fourth the head width (three meas- ured 0.26, 0.26, and 0.27), the median stripe broad, black. Para- frontals silvery but with dark reflections especially opposite antennal insertion; frontals reaching level of arista; parafacials silvery above, grayish below, width at narrowest half that of third antennal joint, about four large bristles and a few hairs on lower part near eye; facial ridges bare except near vibrissae, which are at oral margin; epistoma prominent; palpi black; cheek one-fourth of eye height. Third antennal joint hardly twice the second, arista pubescent near base, its second joint distinct but hardly elongated; back of head not very bulging, with only black hair.
Thorax black with four narrow stripes of white pollen, the outer on the margin, the inner just mesad of dorsocentrals and extending in some lights rather vaguely upon the sides of the scutellum; an- other pair of stripes lie above the supraalar bristles and in rear view extend across the suture anteriorly. Chaetotaxy: dorsocentral 3,3; acrostichal 0,0; humeral 3; presutural 1; notopleural 2; supraalar 2; intraalar 2; postalar 2; sternopleural 2,1; scutellum with two lateral pairs, no apical, the disk with numerous upright slender hairs, no post scutellum. Abdomen black with bluish reflection, viewed from behind with four rows of silvery pollinose spots on second to fourth segments; second segment with one median marginal pair of bristles, third with marginal row of 10, fourth with marginal row of 12 or more; venter black with four pairs of white pollinose spots, not very distinct. First genital segment shining black, with a row of stout bristles; second red with bristles and hairs not in rows on posterior part. Inner forceps red at base, becoming black and slender and curving forward and slightly outward to a sharp tip. Outer forceps
® Proc. U. S. Nat. Mus., vol. 43, 1912, p. 363.
ArT. 1 NEW DIPTERA FROM SOUTH AMERICA—ALDRICH at:
as usual reduced to small red lobes. Penis rather long, black except at base, the tip suddenly bent forward in two flat rounded lobes, somewhat hoodlike, under which several minute sharp processes project toward the base of the organ; about the middle on the front side a flat attachment arises which extends cephalad and then curves broadly toward the tip of the penis, usually resting against it in the spread specimens. Posterior claspers slender and curved clawlike, anterior shorter and less slender, the extreme tip notched into twa minute points.
Wing subhyaline, costal spine distinct, third vein hairy almost to the crossvein, first vein usually with one to four coarse hairs or spinules near middle (absent in both wings in 4 specimens out of 23).
Female.—F¥ ront one-third of head width (0.338, 0.34, 0.37 in three specimens measured); the usual two pairs of orbital bristles. No median marginals on second segment, on the third a noticeable space between the median pair and the rest of the row. Fourth abdominal segment above and three evident sternites below, yellowish red, con- trasting as in the male with the rest of the abdomen. The rows of pollinose spots on dorsal surface of abdomen are larger than in the male, tending to be a little tesselated. The hairs of scutellum and abdomen are less erect than in the male.
Length, 4 to 7.5 mm.
Described from 8 males and 15 females; 20 including the type are from Angol, Chile, collected by D. S. Bullock in December, 1925, and January, 1926; one from Santiago, Chile, collected by A. Faz in 1923; two are from Southern Chile, collected many years ago by M. J. Rivers.
Type.—Male, Cat. No. 40963, U.S.N.M.
The species differs from the type of the genus, Zachina devia Fallen, with which it has been compared, in having the head less bulging behind, the parafacial with stronger bristles, the prescutellars wanting, and the costal spine stronger, as well as in the usual presence of a few setules on the first vein.
Family TACHINIDAE CALLESTHES, new genus
The genotype is a minute muscoid fly with hypopleural bristles, well developed postscutellum and plumose arista. Front rather prominent, antennal axis about one-fifth longer than vibrissal. An- tennae inserted at the middle of the eye; face flat below with very slight keel on upper part; vibrissae at oral margin, which is somewhat above the lower edge of the head. Facial ridges bare; palpi ordi- nary, proboscis small. Cheek nearly one-half the eye height. Male with a single pair of rather large verticals; frontals beginning some
12 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
distance before the ocelli, none reclinate, extending barely to the antennae. Parafacial with delicate hairs in a single row. The dorsocentral immediately before the suture and the last one before the scutellum are much larger than those anterior to them. Abdo- men without discals. Wing somewhat rounded, a well-developed costal spine; first vein bare, rather short, ending just in front of the small crossvein, the large crossvein straight and rather erect, joining the fourth vein half way between the crossvein and the bend, which is obliquely rounded. Apical crossvein concave, the cell ending just
before the apex. CALLESTHES HISTRIO, new species
Male.—The whole fly is opaque black with the following parts white pollinose: the orbits, parafrontal, parafacial, face, a narrow crossband, on the thorax just in front of the suture which extends forward at the sides to the hind edge of the humerus and obliquely downward across the pleura to include the middle of the sternopleura as far as the coxa; another white crossband just in front of the scutellum extending laterally to the postalar calli and anteriorly to the hindmost dorsocentral; the second, third, and fourth abdomi- nal segments have at the front edge a broadly interrupted narrow crossband.
Front 0.21 of head width at the narrowest part, which is a little above its middle, the median stripe is very narrow especially toward the antennae, where it is not more than one-third of the width of one parafrontal. The parafacials are about as wide as the third antennal joint. Antennae black, the third joint one and a half times the second, the extreme base of the arista rather strikingly swollen, suddenly becoming slender where the plumosity begins; palpi black. Cheek one-third the eye height, almost entirely bare, the transverse impression soft and reddish.
Thoracic chaetotaxy: acrostichal 3, 3 (all small); dorsocentral 2, 3; intraalar 2; supraalar 2 (the hind one hairlike); postalar 2; humeral 2; posthumeral 1; notopleural 2; presutural 2 (the inner minute) ; sternopleural 1, 1; scutellum with one large lateral and one large apical, no distinct discal. First abdominal segment with marginal row of bristles mostly rather small; second segment with a marginal row, the median pair and one lateral pair large; third segment with marginal row of eight rather large; fourth segment with a similar marginal row, no discals.
Legs black, pulvilli brown, not much enlarged. Middle tibia with one stout but not long bristle on outer front, two on outer hind side and one flexor. Hind tibia with two bristles only on outer hind side; same on inner hind side.
art. 1 NEW DIPTERA FROM SOUTH AMERICA—ALDRICH 13
Wings dark brown, calypters a little lighter brown; third vein with two hairs at base.
Length, 4 mm.
Described from two males collected by F. X. Williams at Tena, Ecuador, February 17, 1923. The paratype is returned to the Hawaiian Sugar Planters’ Association.
Type.—Male, Cat. No. 40980, U.S.N.M.
The striking and beautiful pattern of the markings is*very similar to Lydellothelaira collaris Townsend and also like Wiedemann’s Zosteromyta dilecta, but these are entirely different in many char-
acters. DAETALEUS, new genus
Belongs to the tribe Theresiini, and agrees with the genotype of Theresia (tandrec Robineau-Desyoidy) in having the fatial keel slight above, disappearing below; palpi and proboscis of ordinary form; no discals on first three abdominal segments and no marginals on first two; and in various minor characters. It differs mainly in having the epistoma much more protuberant, and quite strikingly in possessing the metallic color of many Calliphorids.
Hypopleural bristles and postscutellum well developed. Front not prominent, antennal axis shorter than vibrissal; frontal stripe (male) triangular; only one small pair of verticals, a cluster of proclinate hairs arising from ocellar triangle; frontals barely reach- ing antennae, small; parafacials, eyes and facial ridges bare; antennae arising below level of eye middle, third joint twice the second, arista plumose; vibrissae above epistoma, not approximated ; back of head not bulging. Calypters bare above. Abdomen short and broad, the sternites concealed, only showing a little of first segment.
DAETALEUS PURPUREUS, new species
Male—F¥ront narrow above, before ocelli hardly wider than ocellar triangle and the median stripe reduced to a line; head en- tirely yellow in ground color except upper part of back, and with yellow pollen; palpi and antennae wholly yellow, including arista, which has thin and erect plumosity; beard yellow in most lights but the shorter hairs changeable. Thorax blackish, the sides and scutel- lum metallic. Chaetotaxy: humeral 3, notopleural 2; posthumeral 1; presutural 2 one side, 3 the other in close longitudinal row; acrostichal 0,2; dorsocentral 3,4; intraalar 2; supraalar 3; postalar 2; sternopleural 2, 1; scutellum with 2 large marginals, 1 apical and 1 discal, all represented by scars in the type. Calypters of usual size nearly white. Abdomen wholly metallic purple, mostly
14 PROCEEDINGS OF THE NATIONAL MUSEUM VoL. 74
shining; all the segments with thin whitish pruinosity above in certain angles, more constant below; fourth segment with some smallish bristles, only scars in the type; genitalia (fig. 1) small, with characteristic broad lobe like outer forceps of the “Dexiidae,” the penis long, slender, and jointed. Legs yellow (only the front ones present).
Wings distinctly infuscated; fourth vein obliquely but rather angularly bent, the first posterior cell rather widely open distinctly before the apex; first vein bare, third with three or four small hairs at base.
Length, 8 mm.
Described from one male, collected in Brazil by Beske, sent for identification by the Vienna Natural History Museum, to which it is returned.
Genus INCAMYIA Townsend Fig. 1.—DA®TALEUS PUR- 5 PUREUS. GBPNITAL SEG- Incamyia TownseEnp, Proc. U. S. Nat. Mus., vol.
MENTS AND GENITALIA OF 43, 1912, p. 317. Female only; type and sole MALE FROM LEFT SIDE.
cot ones Sense 3 species, cuzcensis, new.
’ ’ - ’
if, INNER FORCEPS; of, Sphalloglandulus TowNsEnD, Pros. U. S. Nat. OUTER FORCEPS ; cl, CLAS- Mus., vol. 49, 1915, p. 488. Male only; type PER; P, PPNIS. (DRAWN and sole species, S. unicus, new, equals cuz- By C. T. GREENE) censis
The synonomy is from the type specimens in the National Museum. Both species were described from Peru; Townsend placed Jncamyia in the family Phaniidae, and Sphalloglandulus in his Exoristidae. He apparently combined these families with Tachinidae later.
The genus is allied with Lydella and Eucelatoria in having in- frasquamal setules (a small group of minute hairs on the thorax just below the line of attachment of the hind calypter), except in one species noted below, and in having in the female sex a keeled abdomen with piercer of about the same type. From Lydella it differs in having hairy eyes, and from /ucelatoria also by having the facial ridges much more bristly and the second antennal joint longer. Phorocera is also a related genus, but it has the second antennal joint less than half the third, the face more receding, small apical scutellars, and no median upright pair of discal scutellars.
The type species of Zncamyia and the three new species here described form a very compact group occuring in Peru, Chile, and Argentina, all being black in color with mesonotum bearing a median whitish stripe and a well-defined lateral border of the same, so as to give the effect of a heavy black stripe each side of the middle on white ground. The face is nearly vertical, back of head prominent below, cheek equal to nearly one-third of eyeheight. The back of head
art. 1 NEW DIPTERA FROM SOUTH AMERICA—ALDRICH 15
below has mostly black hairs, the white ruff being small. Proboscis and palpi of ordinary form, the latter black or very dark reddish; vibrissae almost at edge of mouth, facial ridges with strong bristles almost meeting the frontals, which descend nearly or quite to the arista. Second antennal joint more than half the third, arista with basal joints short. Female with three pairs of orbital bristles, the anterior proclinate, the other two turned more outwardly; ocellars present and proclinate. Thorax with three posterior dorsocentrals, acrostichals 3,3, sternopleurals 2,1, inner presutural present. Scu- tellum with a median discal pair of upright slender bristles close together, and another pair nearer the tip and farther apart, the space between the discals and the tip bare; three lateral pairs of scutellar bristles, the last long and divergent, no apicals between them. Intermediate abdominal segments with each a single pair of discals. Middle tibia with three or four bristles on outer front side; hind tibia not ciliate, males with front pulvilli and claws moderately elongated. Fourth vein ending far before the tip of wing, third with only a few hairs at base, first bare. Costa with distinct spine.
KBHY TO SPHCIBS OF INCAMYIA
1. First posterior cell petiolate; second to fourth abdominal segments with sharply defined pollinose pale band covering basal third, remainder of these Scoments shining plackis ===) Ss Ses ee ee striata, new species.
First posterior cell open or barely closed in margin_~____----------------- 2
2. Infrasquamal hairs absent; basal pollinose abdominal bands as in preceding,
but still narrower and more or less interrupted in the middle; scutellum
pollinosezonlyat blpeot aes ee ee ee ee spinicosta, new species. Infraquamal hairs present; pollen on abdomen not confined to distinct basal bands; scutellum pollinose from base to tip---___----------------- 3
8. Abdomen mostly gray pollinose, hardly at all tessellated, the pollen in the malerextending upon the venter=22-— 2 =o cuzcensis Townsend. Abdomen mostly dark, the pollen tessellated, in the male not extending upon EG HV EITC Tee era oe Meee aR PA Ve a chilensis, new species.
INCAMYIA SPINICOSTA, new species
Male.—Front one-fourth the headwidth at narrowest part (by micrometer .24, .26 and .26, in three specimens); parafrontal and parafacial silvery, uniform in color, the parafacial a little narrower than width of third antennal joint; cheek one-third the eye height, its anterior part with a few bristles below the transverse impression.
Abdomen shining black, with silvery basal band on segments 2-4; first segment with a large pair of median marginals; second segment with large discal and marginal pairs; third with discal pair and marginal row of 12; fourth with numerous erect discals irregularly arranged. Genitalia small, black. Posterior forceps minute, separate, almost straight; anterior forceps (outer) red at base, flat and blade-hke.
16 PROCEEDINGS OF THE NATIONAL MUSEUM VoL, 74
Wings subhyaline; fourth vein with rounded bend and thence straight to costa well before apex.
Length, 7 mm.
Described from two males from Perales, Chile, September 23, col- lected by Father Anastase Pirion. The place is Quilpue, about 30 kilometers from Valparaiso, These were transmitted by Alfredo Faz of Santiago.
One additional male, collected by Father Pirion, at the same place and on the same day, is larger (8.3 mm.) and has on the middle of the small crossvein in each wing a very large semierect spine, about as long as the hind crossvein. At first I supposed this would be a generic character, but the receipt of additional mate- rial lacking it and agreeing in other respects compels me to believe provisionally that this is a remarkable abnormality of the single specimen.
A female was collected by Edwards at L. Corrientes, Rio Negro Territory, Argentina; it is in the British Museum.
Type.—Cat. No. 28875, U.S.N.M.
INCAMYIA STRIATA, new species
Male-——Slightly smaller than /. spinicosta, differing by the char- acters mentioned in the key. The median pollinose stripe extends very distinctly in its full width to the tip of the scutellum; the basal white pollinose crossbands of the abdomen are very well de- fined and the segments are polished for the greater part of their length. The crossbands end abruptly just under the margin of the abdomen leaving most of the venter shining, or with very slight even pruinosity. Genitalia small, not spread. The apical cross- vein not quite so oblique as in the other species, joining the third vein so as to leave a well defined petiole about as long as the anterior crossvein.
Length, male, 5 and 5.4 mm.
Described from two male specimens collected by C. H. T. Town- send at Oroyo, Peru, May 7 and 28, 1914.
Type.—Male, Cat. No. 40885 U.S.N.M.
INCAMYIA CHILENSIS, new species
Male.—Greatly resembles 7. spinicosta in all respects, but differs in the characters mentioned in the key. When viewed directly from behind the pollen of the abdomen covers thinly almost the en- tire dorsal surface, more dense at the base where in side view it seems to form slightly interrupted crossbands. The fourth segment in side view is shining except at base.
ART, 1 NEW DIPTERA FROM SOUTH AMERICA—ALDRICH td
Female—Front at vortex .34 of headwidth, uniformly widening toward the mouth. Pollen of parafrontal and parafacial with a slight tinge of yellow.
Abdomen with a pronounced keel below on the second and third segments with a good sized curved piercer folded in, the edge of the keel in the third segment bears a series of very short but stout spinules not very noticeable; second segment with a very few spinules almost imperceptible. Dorsally the last three segments of the abdo- men are pretty well covered with pollen which is yellowish on the fourth segment; the tip of the fourth segment and a large indis- tinct subtriangular area on the sides of the second and third are more shining black. The pollen becomes thinner at the margin so that the venter is almost shining.
Length, male 6.4 to 6.7 mm.; female, 5.5 to 6 mm.
Described from eight males and seven females. Four males includ- ing the type, and one female and allotype, are from Angol, Chile, collected in October and December, 1925, by D. S. Bullock; two males and four females from southern Chile (M. J. Rivers), one labeled, “Parasite of Plusia depauperata”; one male from Perales, Chile, September, 1923, collected by Alfredo Faz; one male from Calamarca, Argentina, swept from chrysanthemums by Max Kisliuk, May 238, 1927; one female from La Laoija, Argentina, June, 1927, swept in orchard by Max Kisliuk; and one female from Dolores, Buenos Aires, Argentina, March 29, 1927, swept in orchard by Max Kishliuk.
Four additional specimens are in the Edwards collection from Chile (British Museum)—a male from Llai Llai (a very small village 2 miles from Los Loros), a female from Los Andes, two males from Ancud and Puntra in S. Chiloe.
Shannon collected two males at Los Léros, Chile.
Type.—Male, Cat. No. 40836 U.S.N.M.
REEDIA, new genus
Related to Peleteria, but without palpi and with the bristles very strongly developed, those of scutellum and of abdomen above and below are straight and spine like. Chaetotaxy, head: vertical 2 large pairs; ocellar well developed, proclinate; orbital in female 3 (upper- most smaller); frontals about 11, the upper 2 reclinate and diverg- ing, remainder converging toward middle line, lowest near eye at level of middle of second antennal joint. Thorax: acrostichal 2, 3 (none just before suture) ; dorsocentral 3,4; humeral 7; posthumeral 2; presutural 2; notopleural 2; supraalar 3 very strong; intraalar 3; postalar 2 large and 2 small; sternopleural 2, 1; pteropleural 2 large; scutellum with 2 large laterals and a smaller diverging depressed
18 PROCEEDINGS OF THE NATIONAL MUSEUM VoL, 74
apical pair, besides numerous erect spines. Abdomen: first segment without median marginals; second with about 4 pairs of median marginal spines and some shorter on the disk before them, also 5 pairs at sides; third segment with a marginal row across dorsal surface and continuing below to ends of tergites on the venter, in middle dorsally a few small subdiscal spines as in preceding; fourth segment above and below wholly spiny except anterior third across dorsum. All the sternites bear groups of erect spines.
Proboscis slightly elongate; second antennal joint nearly as long as third, which is convex in front; penultimate joint of arista elongate; face protuberant over mouth, its lateral ridges with only a few bristles above vibrissae.
Type of genus.—Reedia robusta, new species.
REEDIA ROBUSTA, new species
A large, stout, blue-black species, abdomen whollyeshining.
Female—F¥ront broad, 0.36 and 0.38 of the headwidth in the two specimens, subshining bluish-black as far down as the frontal bristles extend; parafacials with silvery white pollen and a few conspicuous black hairs, and bearing two large bristles below (one specimen has two on one side and only one on the other). The ruff of hairs on back of head is deep yellow. Thorax almost destitute of pollen; calypters white. Wings subhyaline; third vein with 8 or 10 hairs at base; bend of fourth vein rectangular; first posterior cell ending in costa far before apex of wing.
Legs black, very bristly; middle tibia with several stout bristles on outer front side.
Length, 15 to 16 mm.
Described from two females collected by Alfredo Faz; one (type) was taken at Concepcion, Chile, the other at Mendoza, Argentina.
Edwards collected two additional females and- Shannon one in Concepcion, Chile; the former are in the British Museum.
Type.—Female, Cat. No. 28896, U.S.N.M.
The genus is named in honor of the late Prof. Edwyn C. Reed, who published several valuable papers on Chilean Diptera, including a catalogue in 1888.
Genus ECHINOPYRRHOSIA Townsend
Echinopyrrhosia TOWNSEND, Ins. Ins. Menst., vol. 2, 1914, p. 90.
ECHINOPYRRHOSIA TROPHOCYON, new species
Differs from the genotype (a/pina Townsend) in having no palpi at all, and in having somewhat more numerous spines on the abdomen. As in alpina, the head is very long, the epistoma remarkably project- ing. Front at vertex 0.31 of the head width, frontal bristles sparse,
ART. 1 NEW DIPTERA FROM SOUTH AMERICA—ALDRICH 19
hairlike above; parafrontals covered with long erect black hairs, which continue on the parafacial. The pollen of the head is dark olive. Parafacial nearly as wide as eye, cheek three-fifths the eye height. Proboscis a little elongated, but not so much as the oral cavity, so it readily folds out of sight entirely. Cilia behind eye remarkably long, back of head with thin ruff of yellow hair.
Thorax blackish, not distinctly striped, the black pile of dorsum long and erect; pleura also with the same pile, especially abundant on mesopleura. Scutellum with dense row of straight, stiff spines on margin and a few upright discal. Calypters dark brown.
ean black without lighter markings, at first seeming to be entirely covered above and below with stout spines; but on hegre til examination the anterior dorsal half or two-fifths of the third and fourth segments and the whole of the very short first segment, except the sides, are seen to be almost wholly bare. There are very abundant spines in the middle of the venter and at the lateral edges, between these regions some shining black bare surface is visible. Femora black, tibiae and tarsi to tip red; claws red basally, pulvili brown. Femora and tibiae very spiny.
Wings entirely infuscated; fourth vein with slightly acute bend a little rounded off, ending far before apex; third vein with only a few hairs at base.
Female.—F ront 0.34 of head width, two proclinate orbitals.
Length, of both male and female, 12 mm.
Described from 1 male and 4 females collected by F. X. Williams, on Mount Tunguragua, Ecuador, January 18, 1923; received from Hawaiian Sugar Planters’ Association, to which two paratypes are returned.
Type—Male, Cat. No. 40982, U.S.N.M.
Genus CHAETOCRANIOPSIS Townsend
Chaetocraniopsis TOWNSEND, Ins. Ins. Menst., vol. 3, 1915, p. 68.
The type and sole original species is C. chilensis Townsend, de- scribed on page 69 of the above reference. It was described from a single male specimen, collected by E. C. Reed in Chile, and now in the United States National Museum. The genus was briefly de- scribed in comparison with Chaetocrania, but the generic characters of the latter were never published, as it was established by the mere citation of a species as type.t| The following notes are taken from the type specimen of chilensis. It resembles the genotype of Gona (capitata De Geer) in having reclinate ocellars, the parafrontals and parafacials wide and with coarse hairs, second joint of arista
4Townsend, Proc. Biol. Soc. Wash., vol. 28, 1915, p. 23; type, Svallanzania antennalis Coquillet.
20 PROCEEDINGS OF THE NATIONAL MUSEUM vou. 74
elongated, face deeply depressed and narrow, its facial ridges high. It differs in having the facial ridges bristly almost up to the arista, which is short and thickened to tip; male without proclinate orbi- tals; the hairs of parafacials and parafrontals much coarser and more erect; the size considerably less, and the body more slender. Otherwise it is structurally very much like Gonia.
KEYS TO SPECIES OF CHAETOCRANIOPSIS
MALES
Arista with blunt tip, pulvilli of front tarsi longer than last tarsal joint; about
13 -pairs! of frontal 2555 = Sas 6 eae eee chilensis Townsend.
Arista with acute tip, although thickened for most of its length; front tarsi
with small pulvilli; about 8 pairs of frontals__--_____ argenticeps, new species. FEMALES
Pollen of parafrontals and parafacials plumbeous__-_----~-~- chilensis Townsend.
Pollen of parafrontals and parafacials silvery__-----_ argenticeps, new species.
CHAETOCRANIOPSIS CHILENSIS Townsend
Chaetocraniopsis chilensis TowNSEND, Ins, Ins. Menst., vol. 3, 1915, p. 69.
Besides the male type, the museum has received two females col- lected by Senor A. Faz at Perales, Chile. They have two pairs of orbitals, the arista is acute at tip, the third antennal joint is one and one-half times the second, the parafrontals and parafacials are somewhat less bristly than in the male. They differ from the females of argenticeps in having the front of head more bristly, as well as in the color of its pollen.
CHAETOCRANIOPSIS ARGENTICEPS, new species
Male.—F¥ront 0.48 of head width at vertex, inner edge of eyes straight and slightly diverging, separated by 0.61 of head width just above vibrissae; parafrontals and parafacials silvery, with less nu- merous bristles than in cAélensis; antennae black, basal joints red, second hardly more than one-fourth as long as third, arista about two-thirds as long as third antennal joint, its apical segment four times the preceding; palpi yellow; cheek one-fourth eye height, dis- tinctly narrower than parafacial, with black hairs; beard of back of head white. Thorax black, with indistinct stripes of pale pollen; dorsocentral 3, 4; sternopleural 4; scutellum with 3 marginal, a spiny suberect apical pair, and a small discal pair; calypters white. Abdomen shining black, basal half to three-forths of second and third segments above white pollinose, which does not continue be- low. First and second segments with a pair of median marginals; third with a stout marginal row of 10; fourth with a smaller mar-
ART, 1 NEW DIPTERA FROM SOUTH AMERICA—ALDRICH DL
ginal row, no discals on any of the segments. Legs black; mid tibia with two erect bristles near middle of outer front side, two on outer hind, and one more sloping on flexor; hind tibia with several irregu- lar on outer hind, two on inner hind and two on outer front side.
Wings subhyaline; third vein with 2-3 setules at base, fourth vein with rounded oblique bend, thence nearly straight to costa, more erect than usual.
Female.—F ront 0.48 of head width at vertex, face 0.58 just above vibrissae. Second antennal joint one-half the third; a pair of proclinate orbitals.
Length, male 7 mm.; female 6.3 mm. }
Described from a pair collected at Perales, Chile, by Rev. A. Pirion.
Type.—Male, Cat. No. 41390, U.S.N.M.
Genus SELENOMYIA Brauer and Bergenstamm
Selenomyia BRAUER and BrErRGENSTAMM, Zweifl. Kais. Mus., pt. 5, 1891, p. 861; pt. 6, 1893, p. 170—AtpricH, Annals Ent. Soe. Amer., vol. 18, 1925, p. 459.
The type species, and the only one so far known, is S. brevicornis Brauer and Bergenstamm, mentioned and more or less described in all the above references. Only cone specimen is known, the type, which I borrowed from the Vienna Museum and later sent back. I am now describing three additional species which I refer to this genus; like the genotype, they are from Chile, where it seems to be a characteristic form. The third species, facialis, differs from the others in having the parafacials hairy and may ultimately go into a new genus; but from the slight taxonomic value of this character in other dexiid groups, as, for instance, Ptilodexia and FRutilia, I do not believe it is generic in the present case.
In this genus there is a high, sharp carina on the face, and the facial ridges are decidedly convergent and prominent below, so that there are well-marked and deep subantennal depressions which are entirely separated. The cheek is almost equal to the eye height and the parafacials are broad, about two-thirds of the cheek.
KEY TO THE SPECIES OF SELENOMYIA
1. Abdomen shining green, without pollen_____-___________~_ virens, new species. Abdomen plack-awith pollinose markines!=)) 2 2
2. Tibiae reddish-yellow ; second and third abdominal segments with a pair of arcuate spots of pollen_____________ brevicornis Brauer and Bergenstamm. ibiae black, abdominal’ spots*notsarcuates 92s s ees ee ee 3
3. Parafacials bare except on upper part_--_________-_____ plena, new species.
Parafacials with coarse hair down to the lower edge of the eye. facialis, new species.
22 PROCEEDINGS OF THE NATIONAL MUSEUM VoL. 74 SELENOMYIA VIRENS, new species
Male.——Dark metallic green in color, the abdomen wholly shining. Arista bare, hypopleural bristles and postscutellum well developed. Front at vertex narrow, only 0.16 of head width. Dimensions of head in micronometer units: height 66 (eye 38); length (antennal axis) 48; width 74. Vibrissal axis almost as long as antennal, but both back and front of head recede a little below. Pollen of para- frontals dark gray, that of parafacials decidedly brown, between is a large blackish spot on each side of the base of antennae, most dis- tinct from in front. Vibrissae about two-thirds the length of the third antennal joint above the oral margin; facial ridges densely bristly and decidedly convergent on lowest third; face with thin, sharp carina, forming decided antennal grooves; antennae black, third joint hardly twice the second, arista of moderate length, its penultimate joint distinct but hardly elongated. Palpi dark yellow, of normal size; proboscis short, fleshy. Beard black.
Thorax dark green, with three broad stripes of white pollen which are distinct at front edge, but fade on the disk and are hardly visible at the suture. Humeri and sides of thorax before wings also with white pollen. Chaetotaxy: acrostichal 3, 8; dorsocentral 3, 4; hu- meral 6; posthumeral 3; presutural 1; notopleural 2 (with hairs around bases) ; supraalar 3-4; intraalar 3; postalar 2; sternopleural 2, 1; scutellum with three large lateral pairs, of which the third might be called large divergent apicals, and one or two pairs of widely spaced smallish discals. Calypters brown with blackish border.
Abdomen wholly shining dark green; second abdominal segment with two pairs of median marginals and two pairs of discals, one behind the other; third segment with two pairs of discals and a mar- ginal row; fourth segment with a few erect bristles irregularly placed. Fifth sternite deeply cleft, the lobes with hairs only; genital segments small, dark brown; inner forceps united into a slender, almost needlelike organ, the outer forming broad concave plates as in many dexiids, but in this species decidedly truncate; penis a very slender black plate with a pale prolongation and at the base a pair of erect bladelike semitransparent pieces (claspers?).
Wings subhyaline, first posterior cell open and ending considerably before the apex; fourth vein curved backward at the bend, which is rectangular and slightly rounded; third vein with three or four setules at base.
Legs black, claws and pulvilli elongated, the latter brownish; front tibia with two outer bristles; middle tibia with three on outer front side; hind with a few irregular on outer side.
ART. 1 NEW DIPTERA FROM SOUTH AMERICA—-ALDRICH 2a
Female —F¥ront 0.27 of head width; two pairs of proclinate orbi- tals. Second abdominal segment with a single pair of median marginals, no discals.
Described from two males and one female, collected at Perales, Chile, near Santiago, by Rev. Anastase Pirion.
Type.—Male, Cat. No. 41382, U.S.N.M.
SELENOMYIA PLENA, new species
Male.—Like virens, but lacking the green color, and with abdomi- nal markings. Black throughout, only the palpi yellow; parafacials brown, but without distinct blackish spot above; calypters white, the rim infuscated. Chaetotaxy as in virens, but the inner presutural is present; the third antennal joint is bulbous at tip; the dorsal pollinose stripes of the thorax are more distinct, being visible almost to scutellum; second and third abdominal segments with white pollen except on a broad median shining stripe and the posterior third of each, this pollen extending underneath almost to the middle. Fourth segment with very thin pollen on same portions. Second segment with one discal and one median marginal pair, third with one discal pair and marginal row. Gentalia black; the united inner forceps awl- shaped, the tip blunt, rounded, and turned back; outer forceps slender and pointed, not platelike. Front 0.24 of head width.
Described from one male, collected by D. S. Bullock at Angol, Chile, January 2, 1927; and a male and two females collected by Edwards at Concepcion, Chile; Peulla, S. Chile; and Lake Nahuel Huapi, Rio Negro, Argentina.
Type.—Male, Cat. No. 41383, U.S.N.M.
SELENOMYIA FACIALIS, new species
Female——F¥ront 0.26 of head width; only one pair of proclinate orbitals, which are high up and almost in the frontal row; the upper- most frontal small and turned outward (it is broken off in the female of virens, so I can not compare the direction and no other females are known) ; parafrontal and parafacial gray-brown pollinose, with- out distinct spot at antennae. Palpi brown; facial ridges on the convergent lower part not so bristly as in virens,; arista pubescent; beard pale yellow except anteriorly; chaetotaxy and thoracic stripes as in virens. Abdomen black, almost covered with thin gray pollen, subshining in some lights, hind edges of segments shining; second segment with a marginal pair, third with marginal row, both with only a few stouter hairs in place of discals. Calypters white, the rim infuscated only on inner side. Legs as in virens, but the front tibia has a single outer bristle, and the hind one has a trace of reddish color on the middle. Wing as in wrens.
24 PROCEEDINGS OF THE NATIONAL MUSEUM VOL, T+
Described from one female collected at Perales, Chile, by Rey. Anastase Pirion. Type.—Female, Cat. No. 41384, U.S.N.M.
PIRIONA, new genus
Hypopleural bristles and postscutellum well developed; ocellars large in both sexes, curving directly to the side, neither proclinate nor reclinate; eyes and parafacials densely pilose; frontals ending hardly below base of second antennal joint; outer verticals not devel- oped, inner strong, just behind them on occiput a smaller pair of con- vergent bristles, almost decussate; back of head bulging, front not prominent, antennal axis slightly exceeding vibrissal; cheek one-third of eye height, parafacial one-half of cheek; facial ridges bare, not prominent, vibrissae at oral margin, which is slightly above lower edge of head, not protruding; palpi normal, proboscis short, fleshy ; anten- nae inserted just below level of middle of eye, reaching nearly to vibrissae, third joint rather wide, hardly twice as broad as long and much less than twice the second; arista bare, penultimate joint short; female with usual two pairs of proclinate orbitals. Scutellum with two laterals, a long decussate apical pair, and several irregular erect straight bristles. No acrostichals immediately before suture. A few minute infrasquamal setules present. Abdomen without median mar- ginals on first segment, a pair of large discals and a pair of large marginals on second, third with a pair of large discals and a marginal row (marginals of second and third rather far forward). Female without piercer. Legs with large bristles, middle tibia with one large on flexor side, hind with several large bristles widely spaced on outer hind side. Wing of ordinary form, no costal spine, first vein bare, third with a few large hairs at base; fourth vein with angular, slightly oblique bend, ending distinctly before tip, the hind cross vein straight and joining fourth vein at last fourth of distance from small cross vein to bend.
I am unable to find a genus in the national collection with which this is closely related.
Genotype.—Piriona fasciculata, new species.
PIRIONA FASCICULATA, new species
Male—Black, palpi at least basally and the tip of the second an- tennal joint reddish yellow. Front wide, 0.29 of head width on upper third, widening gradually ; frontals 8, decussate, none reclinate, the first anterior to the ocellar triangle; frontal stripe wide, black; parafacials and parafrontals black, subshining; beard black. Thorax black, shining, with traces of white pollen at front edge only. Chaetotaxy: acrostichal 2, 2 (hindmost large and rather far before
anv. 1 NEW DIPTERA FROM SOUTH AMERICA—ALDRICH 25
scutellum) ; dorsocentral 3, 8; humeral 4; posthumeral 1; presutural 1; notopleural 2 (surrounded by hairs); supraalar 3; intraalar 3; postalar 2; sternopleural 2, 1; scutellum as mentioned. Calypters white, the rim blackish on inner edge. Abdomen broad and rather deep, the genital segments giving it a truncate appearance, sub- shining black but the intermediate segments with some thin white pollen in oblique view; bristles of fourth segment large and erect anteriorly, becoming denser and smaller on posterior part. First genital segment rather
prominent, declivous, o second more _ bulging. \ Sy
Inner forceps united into a slender black or- gan slightly bent for- ward at tip; outer for-
ceps also black and slen- Fic. 2.—PIRIONA FASCICULATA, NEW SPECIES. HIND LEG OF MALE. (DRAWN BY C. T. GREENE)
der; penis and claspers black but not distinctly visible in the only specimen. Legs rather stout, front claws and pulvilli a little elongated; mid tibia with several bristles on outer front side, one at the middle fully half as iong as the tibia, and one of almost the same length on the flexor side; hind tibia besides those on outer hind side has three on outer front and the same on inner hind. Hind coxa with a fasciculus of long black hairs on inner edge; hind femur curved, with a prominence at two-thirds of its length below, bearing a remarkable fasciculus of long black hairs. (Fig. 2.) Wing grayish, gradually becoming more clouded toward base.
Female.—Front at vertex 0.80 of head width, very gradually wider anteriorly. Genital segments not in position to examine, but evidently without piercer. Hind femur without fasciculi.
Length, 6.2 mm.
Described from one male and one female collected at Marga Marga, Chile, by Rev. Anastase Pirion, after whom I name this remarkable genus. Also from one male collected by Edwards at Bariloche, Rio Negro, Argentina (British Museum) ; and three males collected by Shannon, two at the place just mentioned, the other at Lago Gutierrez, Rio Negro, Argentina.
Type—Male, Cat. No. 41385, U.S.N.M.
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FHREE NEW SPECIES OF TWO-WINGED FLIES OF THE FAMILY BOMBYLIIDAE FROM INDIA
By J. M. Avpricw
Associate Curator, Division of Insects, United Siates National Musewm
The following new species of Bombyliidae were reared by C. P. Clausen as secondary parasites on Hymenoptera of the genus Tiphia, which were parasites of scarabaeid beetles of the subfamily Rutelinae in India. Mr. Clausen also reared Hyperalonia oenomaus Rondani with the same habits and in the same region.
APHEOGEBANTUS CLAUSENI, new species
Male—Ground color black except femora and tibiae. Head glo- bose, hind margin of eye with the usual deep notch and bisection ; occiput with deep cleft behind vertex; front at vertex twice as wide - as ocellar triangle, gradually widening to antennae, where it is about one-fourth of the head width; face receding; front and face with black hairs, the former with golden appressed scales, the latter with erect yellow hairs mixed with the black; antennae short, the third joint obliquely onion-shaped, the basal enlarged part shorter than second joint, the style about equal to half the slender part of the joint. Dorsum of thorax and scutellum thinly covered with ap- pressed deep golden tangled hairs mixed with erect slender black ones; a collar of yellow hairs next to head; pleurae glaucous, meso- pleura with abundant yellow hairs; three rather large black bristles before base of wing and several on postalar callus; scutellum with four pairs of black bristles on margin, equally spaced. Abdomen with same golden appressed hair as mesonotum, mixed with black hairs not so erect; first segment with dense brush on each side of erect blunt yellow hairs. Legs yellow, the coxae, base of front femora, tip of hind femora above, and tarsi black: all the tibiae with rows of small spines; pulvilli normal; hind femur without row of bristles on lower hind edge. Halteres yellow, distal part of stem and base of knob infuscated. Wings brown, gradually paler poste- riorly; second vein branching from third opposite extreme base of
No. 2747.—PROCEEDINGS U. S. NATIONAL Museum, VoL. 74, ART. 2 2606—28 1
2 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. T4
diseal cell; anal cell wide open; second vein joining costa at a right angle.
Length, 4.5 to 6 mm.
Described from three males, Shillong, India.
Type.—Male, Cat. No. 40301, U.S.N.M.; one paratype will be de- posited in the British Museum.
Only one species of Aphoebantus has been described from India heretofore, ceylonicus Brunetti, which has the thoracic and scutellar bristles yellow, as well as the erect: hairs on thorax and abdomen; the description indicates that the abdomen is more or less banded, which is not the case in A. clausend.
APHOEBANTUS SERATUS, new species
Male.—Differs from A. clauseni only in the following: The front at vertex is considerably narrower; the middle femora are black on basal half or more; the hind femora have a row of distinct spines on iower inner edge from base almost to tip; the wings are paler, the only distinct infuscation being at extreme base and in the subcostal cell; the first and third veins on the dorsal surface are smooth, while under high power they appear dotted in A. clauseni as if hairy, although I can see no hairs on them in my specimens. Third _antennal joint missing.
Length, 5.2 mm. Described from one male, Shillong, India. Type.—Male, Cat. No. 40302, U.S.N.M.
EXOPROSOPA SIPHO, new species
Female—Ground color black except the tibiae and_basitarsi. Wings deeply infuscated on basicostal half with spots on the forks of veins in the clear. portion.
Front of head at vertex about one-fifth the width of the head, widening to one-third at the antennae. Thinly sprinkled with elongate yellow appressed scales and erect black hairs. Face with the same covering, slightly brown at edge of mouth. Proboscis con- siderably elongate, when directed forward exceeding the antennae.
Thorax rather sparsely sprinkled with the same scales as on the head, more dense on scutellum; some black bristles in front of wing and on the postalar region, the latter also having rather dense and somewhat elongated yellow hair. Mesonotum and scutellum also with erect delicate hairs which are more brown than black in the middle part. Pleura with a dense tuft of black hairs on upper half of mesopleura, the lower half with thin and delicate but rather long hair. Sternopleura with a few mostly yellowish hairs which extend upward along the anterior part of the pteropleura, the metapleura with long dense bristly hairs which are about one-half yellow.
ART. 2 THREE NEW SPECIES OF TWO-WINGED FLIES—ALDRICH 3
Abdomen black with mixed black and yellow appressed scales. On the first segment there are dense long, erect, scaly hairs at the sides, continuing in a thin and reduced series across the middle. The second segment shows a few paler scales across the base, hardly enough to call crossbands; the following segments show scarcely any banding except the last three which when viewed from behind show mostly black scales across the base. The sides of the abdomen are fringed with large blunt, black scales, a few whitish. The venter is covered with long, slender, yellow hairs not very dense. Legs cov- ered with mixed light and dark scales. Pulvilli wanting; hind femur with a row of about seven bristles on the lower front edge. Wing dark brown on basicostal half, brownish hyaline on the remainder, the line of division not very distinct, running obliquely from the mid- dle of the axillary cell to the costa just before the apex. All the forks and crossveins bear spots and there is a slight trace of addi- tional infuscation along the anterior fork of the second vein near its tip, a less distinct trace on the posterior fork of the same and a slight spot in the open apex of the anal cell.
Length, 4.6 and 4.8 mm.
Described from two females, Shillong, India.
Type—FKemale, Cat. No. 40303, U.S.N.M.
This species is close to Hwoprosopa insulata Walker, but ac- cording to the descriptions of Brunetti and Nurse /. sipho differs in having a much longer proboscis, the dense bristly hairs of the pleura confined to the upper edge, the abdomen with very indis- tinct bands and with well-developed scales along the edges, and the venter covered with pale hairs. In the paratype specimen the proboscis would, if directed forward, extend a little beyond the antennae, but in the type the organ is lengthened out as if by the straightening of an elbow so that it is about as long as the whole body not counting the head; or as it projects downward from the head its length is equal to about two and a half times the vertical diameter of the head. The paratype specimen will be deposited in the British Museum.
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NEW FOSSIL PEARLY FRESH-WATER MUSSELS FROM DEPOSITS ON THE UPPER AMAZON OF PERU
By Wiiu1am B. MarsHaty Assistant Curator, Division of Mollusks, United States National Museum
The United States National Museum has recently received some fossil Naiads collected by Prof. Joseph T. Singewald, of Johns Hopkins University, in deposits at the headwaters of the Upper Amazon, Peru; also another sending collected by Dr. Harvey Bassler, likewise from Peruvian deposits. The exact geological horizon from which these shells were obtained has not been definitely settled to date. Conrad? says, “It is not possible to state without doubt what the relative stratigraphical position of this group may be, but if all the species are extinct it can not be later than the Tertiary. The Pleistocene origin of the group is at least very doubtful.” None of the species found in these deposits are living to-day.
Associated with these mussels were large numbers of Anisothyris, Isaea, Neritina, and other fossils, which are brackish-water forms. It is quite possible that the Naiads were swept in to these deposits from higher levels by floods, and thus came to be mingled with the brackish-water faunas. These shells, therefore, argue for the existence of bodies of fresh water at the time they were living at or near the place where they were found.
Until now only one Naiad has been described from the deposit, namely, Anodon batesiti Woodward. As Anodon (=Anodonta) does not occur in South America, this shell is probably an Anodontites belonging in the family Mutelidae. The new species described herein belong in the subfamily Hyriinae of the family Mutelidae and are the first fossil species of that subfamily described from South America. They are related to two genera, Diplodon and Hyria, as will be shown in the remarks under the new genus Prodiplodon.
PRODIPLODON, new genus
Pearly fresh-water mussels of the family Mutelidae with beak sculpture resembling that of the genus Diplodon and also that of
1 Amer. Journ. Conch., vol. 6, p. 192, 1870.
No. 2748.—PROCEEDINGS U. S. NATIONAL Museum, VoL. 74, ArT. 3. 2607—28 1
> PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
the genus Hyria, and consisting of several V-shaped undulations, “nesting” one within the other, and other undulations on the ante- rior and posterior umbonal areas, each pair of which if continued would form another V in the series.
Type of the genus—Prodiplodon singewaldi, new species, de- scribed in this paper. |
This genus is distinctly related to Diplodon and to Hyria and seems to stand about midway between them. In Diplodon the undu- lations are more nearly direct radial than V-shaped. In Hyria they are composed of “nested” V’s but in that genus the shell is alate anteriorly and posteriorly. The fossils at hand give no indication of having been alate posteriorly, but in the type the anterior end is minutely winged. It is possible that this genus was the forerunner of both Hyria and Diplodon. Prodiplodon may have one living representative in “ Castalia” pazi Hidalgo,? the type locality of which is Imbabura, Ecuador. Simpson* places it in the genus ‘Diplodon. From the description and figure it appears to be a ‘Prodiplodon and a direct descendent of the new fossil species Prodi- plodon singewaldi. It appears to be the only Diplodon-like species living in the region near the localities in which the fossils were found. True Diplodon is most plentiful in the La Plata system and in south- eastern Brazil. It occurs also in Chile and eastern Brazil, a couple of species in Guiana, and a couple in Patagonia. The abundance of ‘Diplodon, evidently a descendent of Prodiplodon, in the La Plata system is interesting when considered in connection with Hrodona (apparently descended from Aznisothyris, the most plentiful fossil in the deposits under consideration) which is found in the brackish waters of the La Plata estuary.
PRODIPLODON SINGEWALDI, new species Plate 1, figs. 3, 6
Shell rather compressed, somewhat quadrate in form, rounded in front, slopingly truncated at the rear. Ventral margin a little curved in its median portion, sweepingly rounding into the anterior margin and making a broad angle with the posterior margin. Dorsal margin arched, making a very broad, scarcely noticeable, angle with the posterior margin. Beaks well forward, about 19 mm. behind the extreme anterior end and 51 mm. in front of the extreme posterior end. Posterior ridge low, slightly angulated at upper part of shell, anterior ridge almost entirely lacking. Sculpture of very prominent concentric lines marking rest periods, with minor concentric striae be-
tween them. Umbonal area sharply and closely sculptured with ra- eT gg eee 2 Journ. de Conch., vol. 8, p. 353, pl. 13, fig. 6, 1868. 8 Descriptive Cat. Naiades, p. 1273, 1914.
ART. 3 NEW FOSSIL MUSSELS FROM PERU—MARSHALL aS
diating threads or riblets. Those at the anterior portion are narrow, curved, somewhat granulous and do not rapidly increase in thick- ness. Those at the posterior end are also curved but increase rapidly in size. In the middle portion the threads join into several! V’s with nearly straight sides. The curved threads of the front and rear por- tions if continued would form V’s with curving sides. Some of the sculpture shows a tendency to continue into the next area after the distinct umbonal period was passed; at the anterior end this tendency shows itself in several direct but rather obscure threads which con- tinue across the next growth area. Pseudocardinal teeth plate-like, nearly parallel to the dorsal margin and close to the tip of the beak. Lateral tooth long and curved.
The type (Cat. No. 370808, U.S. N. M.) measures: Length, 70 mm. ; height, 39 mm.; diameter, if both valves were present, would be about 20 mm. A paratype forms Cat. No. 370809, and a paratype was returned to Professor Singewald. They came from Paucarpata, on the Maranon River, and were collected by Dr. Joseph T. Singewald, jr., in whose honor the species has been named. The most striking feature of this species is the neat, clean-cut character of the umbonal sculpture. The species is related to Prodiplodon paucarpatensis described in this paper, but is less elongate and somewhat more nasute. It is related also to Prodiplodon basslert Marshall which, however, has an elliptic outline and the beak sculpture pointing differently. The three species form a very natural group.
PRODIPLODON BASSLERI, new species
Plate 1, fig. 1
Shell moderately thin, rather compressed, elliptic, slightly nar- rower in front. Ventral margin regularly curved, rounding regu- larly into the posterior margin, and apparently into the anterior margin also. Posterior ridge low and rounded. Anterior ridge not differentiated from the general surface. Beak set well forward, about 10 mm. behind the anterior margin, and 50 mm. in front of the posterior margin. Sculpture of fairly well-marked concentric striae and lines showing rest periods—the earlier ones set obliquely across the general surface of the shell. Sculpture of umbonal area somewhat obscured but evidently similar to that of Prodiplodon singewaldé but the points of the V’s pointing in the general direc- tion of the postero-ventral margin, while in Prodiplodon singewaldi they are directed toward a point anterior to the middle of the ven- tral margin. Pseudocardinal teeth placed directly under the tip of the beak. Anterior adductor scar very deep, posterior scar nearly superficial. Pallial line at rear end about 10 mm. from the ventral margin.
4 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
The type, (Cat. No. 370811, U.S.N.M.) measures: Length, 60 mm.; height, about 38 mm.; diameter, if both valves were present, would be about 16 mm.° It was collected at Pebas, Peru, by Dr. Harvey Bassler in whose honor the species is named. Related to Prodiplodon singewaldi described in this paper, from which it dif- fers in having the outline elliptic instead of quadrate. It groups also with Prodiplodon paucarpatensis.
PRODIPLODON PAUCARPATENSIS, new species Plate 1, fig. 4
Shell compressed, elongate trapezoidal, slightly narrower in front, slopingly truncated at the rear. Dorsal margin nearly straight, ventral margin nearly straight, rounding into the anterior margin, Joining the posterior margin at an angle. Beaks well for- ward, about 18 mm. from extreme anterior end, 56 mm. from ex- treme posterior end. Descent from beaks to anterior margin rather abrupt. Posterior ridge low and rounded. Posterior dorsal area long and broad. Sculpture of well-marked concentric lines of growth. Umbonal portion of shell with sculpture evidently similar in general pattern to that of Diplodon singewaldi described in this paper, but weathered, so that details can not be pointed out. The whole shell lacks its outer surface and is nearly white.
The type (Cat. No. 370810, U.S. N.M.) measures: Length, 74 mm.; height, 39 mm.; diameter, 18 mm. It comes from Paucarpata, on the Maranon River, and was collected by Dr. Joseph T. Singewald, jr., of Johns Hopkins University. This species has the form com- mon to the well-known Unio complanatus of the United States. It is related to Prodiplodon singewaldi Marshall, but differs in form, being more elongate and less oblique. It is related also but not so closely to Prodiplodin bassleri.
EOLRIPLODPGN, new genus
Beaks with very coarse, nearly direct radial undulation, some of which are broken up into nodules. Close to the tip of the beak each pair of undulations unites into a V but the later ones become nearly direct and do not form a V.
Type of the genus, Lodiplodon gardnerae Marshall, described below. This genus also is related to the recent genus Diplodon.
EODIPLODON GARDNERAE, new species Plate 1, figs. 2, 8
Shell rather thin, very elongate, beak set very far forward, about 7 mm. behind extreme anterior margin, 57 mm. in front extreme posterior margin. Dorsal margin somewhat broken but evidently
ART. 3 NEW FOSSIL MUSSELS FROM PERU—-MARSHALL 5
nearly straight posterior to the beak and sloping downward to the posterior margin. Anterior to the beak the very short dorsal de- scends rapidly to meet the anterior margin. Ventral margin nearly regularly curved, sweeping into the anterior margin in a broad curve and not differentiated from it. Posteriorly the shell becomes nar- rower, the ventral margin curving sharply to join the posterior mayr- gin, the two margins differentiated from each other by the end of the posterior ridge. Posterior ridge well marked, with an indistinct rib running along its summit. Posterior area with two faint radiat- ing grooves forming a low, broad rib between them. General surface of shell rather smooth, concentric growth lines not well marked except on the posterior area and near the anterior margin. At the anterior end of ventral margin are five short radiating flutings. A faint groove just appreciable to touch and made faintly visible by a dark line extends from the beak to the ventral margin just behind its middle point, probably marking the depth to which the animal in- serted itself into the material of the bottom. Interior pearly, ante- rior adductor scar deep, posterior scar well marked but shallow. Pallial line well marked, about 6 mm. from the ventral margin. Umbonal area very prominent because of the strong radial sculptur- ing. At the anterior end this sculpture consists of five fine, rather granular, curving sharp threads, the middle area shows a crudely V-shaped arrangement, partly broken up into stout nodules. The posterior portion has several direct radiating threads or riblets.
The type (Cat. No. 370812, U.S.N.M.) measures: Length, 64 mm. ; height, about 30 mm.; diameter if both valves were present would be about 16mm. It was collected by Doctor Singewald at Pebas, Peru, and is named in honor of Dr. Julia A. Gardner, of the United States Geological Survey. Cat. No. 370814 includes a cast, a partially preserved umbonal portion, and a number of fragments, all from the type locality. Several partially preserved specimens were re- turned to Professor Singewald.
The type, owing to a rich chestnut coloring due to a ferruginous stain, has almost the appearance of a recent shell. The species is not closely related to any known recent species. In boldness and extent of umbonal sculpture it is scarcely equaled by any recent spe- cies, and in the breaking up of part of the sculpture into nodules it stands unique.
EODIPLODON PEBASENSIS, new species
Plate 1, figs. 5, 7
Shell apparently rather compressed, thick, rounded in front, obliquely truncated behind. Dorsal margin lightly arched, ventral margin slightly curved, meeting the posterior margin in an obtuse angle, and rounding into the anterior margin. Posterior ridge low
6 PROCEEDINGS OF THE NATIONAL .MUSEUM VOL. 74
and not well pronounced. Beak set well forward, about’ 5 mm. from anterior edge and 20 mm. from posterior end. Concentric sculpture of faint growth lines. Umbonal region boldly sculptured with radiating ribs and nodules. The anterior portion has five direct, sharp, faintly granulate radiating threads. The middle portion has the riblets arranged in V pattern and somewhat nodulous. The pos- terior portion has four sharp, direct slender threads. Pseudocardinal tooth stout.
The type (Cat. No. 370813, U.S.N.M.) is:a fragment but includes all the shell up to a well-marked rest period. At that period it meas- ured: Length, 25 mm.; height, 13 mm. It came from Pebas, Peru, and was collected by Dr. Joseph T. Singewald.
In beak sculpture this species resembles and might be mistaken for Hodiplodon gardnerae, but careful examination shows marked differences. In form the two species are very different, Hodiplodon gardnerae being very elongate, narrow, and rather inflated, while Eodiplodon pebasensis is subquadrate, broad, especially at the pos- terior end, and rather compressed.
ANODONTITES?
Included with the collection sent by Doctor Singewald are some fragments of a very large fossil pearly fresh-water mussel, from Tarapoto, Peru. Cat. No. 370815. There is not enough left to determine the genus, but what we know of the microscopic structure of the South American Naiades leads to the belief that they belong to some genus in the subfamily Mutelinae of the family Mutelidae. The shell must have been very large and massive, as one of the fragments from along the ventral margin has a thickness of about 8 mm., and the pallial line is about 27 mm. from the ventral edge. These measurements are not equaled by any known South American Naiad, recent or fossil. They call to mind the size and massive- ness of our largest North American Naiades, namely, the genus Crenodonta. The numerous layers of nacreous material, each very thin, between the pallial line and ventral margin indicate that the shell was aged, and that growth at this period was very slow. Dur- ing this time the pallial line appears to have remained nearly sta- tionary. The prismatic layer is very thick (in some places about a millimeter) indicating a member of the Mutelinae and not the Hyriinae, as in the latter the layer is thin and usually flakes off with the periostracum. In spots the component spicules of the prismatic layer have separated from each other and lie scattered about or in a heap.
ART. 3 NEW FOSSIL MUSSELS FROM PERU—MARSHALL a
BIBLIOGRAPHY
The following list of papers comprises about all that has been published on the formations considered in this paper:
1869. W. M. Gass, Descriptions of fossils from the clay deposits of the Upper Amazon, Amer. Journ. Conch., vol. 4, pp. 197-200, pl. 16.
1871. T. A. CoNRap, Descriptions of new fossil shells of the Upper Amazon, Amer. Journ. Conch., vol. 6, pp, 192-198, pl. 10.
1872. C. F. Hartt, On the Tertiary basin of the Maranon, Amer. Journ. Sci. and Arts, ser. 3, vol. 4, pp. 58-58.
1878. Oskar Bortreer, Die Tertiarfauna von Pebas am oberen Maranon, k. k. Reichsanstalt, vol. 28, pp. 485-504, pls. 13 and 14.
1879. C. BARRINGTON Brown, On the Tertiary deposits on the Solimoes and Javary Rivers in Brazil, Quar. Journ. Geol. Soc. London, vol. 35, pp. 77-82.
1879. R. Erueriper. Note on the mollusca collected by C. Barrington Brown from the Tertiary deposits of Solimoes and Javery Rivers, Brazil. Quar. Journ. Geol. Soc. London, vol. 35, pp. 82-88, pl. 7. This paper is an appendix to the preceding one. ;
1921. M. G. pe OttvimrA. Breve noticia sobre os fosseis terciarios do Alto Amazonas, Boletin Service geologico e mineralogico do Brasil. No. 11, pp. 41-52, 1 plate.
1927. JuLIA GaRpNER. A recent collection of late Pliocene invertebrates from the headwaters of the Amazon, Journ. Washington Acad. Sci., vol. 17, no. 20, pp. 505-509.
EXPLANATION OF PLATE
Fig. 1. Prodiplodon bassleri, new species nat. size.
. Hodiplodon gardnerae, new species nat. size.
. Prodiplodon singewaldi, new species nat. size.
. Prodiplodon paucarpatensis, new species nat. size.
. Hodiplodon pebasensis, new species nat. size.
. Prodiplodon singewaldi, new species. Beak sculpture X 2 diameters. . Hodiplodon pebasensis, new species. Beak sculpture X 2 diameters. . Hodiplodon gardnerae, new species. Beak sculpture X 2 diameters.
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VOL. 74,
PROCEEDINGS,
NATIONAL MUSEUM
U. S.
WATER MUSSELS FROM PERU
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TWO NEW NEMATODE WORMS FROM RODENTS
By Emmertr W. Price
Of the Zoological Division, Bureau of Animal Indusiry, United States Depart- . ment of Agriculture
In this paper two nematodes which appear to be new species from rodents are described. ‘The first of these species was collected from the cecum and colon of a flying squirrel, Glaucomys volans volans, which was found dead by Miss Florence Thompson, librarian of the Bureau of Animal Industry, at her home at Falls Church, Va., July 28, 1927. 'This parasite belongs to the family Oxyuridae Cobbold, 1864, subfamily Syphaciinae Railliet, 1916, and genus Syphacia Seurat, 1916. For this species the name Syphacia thompsoni is proposed.
The second species was collected from the small intestine of a gray squirrel, Scturus carolinensis, which was killed near Bowie, Md., by Dr. Albert Hassall, October 10, 1927. This species belongs to the family Trichostrongylidae Leiper, 1912, subfamily Helimosominae Travassos, 1914, and genus Heligmostrongylus Travassos, 1917. For this species the name Heligmostrongylus hassalli is proposed.
SYPHACiIA THOMPSONI, new species
Specific diagnosis —Sy phacia: Slender forms, milk white in color, male much smailer than female, and with the tail of both sexes very long and slender. Cuticula coarsely striated transversely. ‘The mouth is provided with three lps of about equal size. The cir- cumoral papillae and amphids are situated laterally in two groups, each group consisting of two small submedian papillae and a large umbilicated amphid or so-called lateral papilla. (Fig. 1.)
Male 3.1 mm. long and 156 to 160% wide. The posterior end of the male is strongly curved ventrally in the form of a hook. (ig. 2.) The cuticle of the anterior end of the body is inflated and is 78 to 98 in diameter and extends from the base of the lips to the level of the cervical papillae. The esophagus is club-shaped, 250 to 280 long by 388 wide at the narrowest part and increasing to 60, at the posterior end. The esophageal bulb is spherical, 76 to 994 in diam-
No. 2749.—PrRocEEDINGS U. S. NATIONAL MUSEUM, VOL. 74, ART. 4 2608—28 1
2 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
eter, and is joined to the esophagus by a short slender constriction. The nerve ring surrounds the esophagus about 90 from the anterior end of the body. The excretory pore opens ventrally 75u caudad of the esophageal bulb. ‘The cervical papillae are situated 172 to 214y from the anterior end of the body. The lateral alae arise imme- diately caudad of the cervical papillae and extend to within a short distance from the cloacal aperture.’ The tail is long and slender, and terminates in a fine point. (Fig. 3.) There are two narrow, sym- metrical bursal alae and three pairs of caudal papillae, one pair pre- anal, one pair adanal, and one pair postanal; the postanal papillae are large and pedunculated, and support the caudal alae. The clo- acal aperture is situated on a slight prominence 340 to 4354 from the end of the tail. Immediately caudad of the cloacal aperture there is a small rounded prominence which bears six small spikelike projections. The spicule is simple, slightly curved, and 156 to 190p long by 18 wide at the base. The gubernaculum is 95 to 110, long, directed transversely, and is provided at its distal end with a barb- like hook. (Fig. 4.) The ventral surface of the body bears three large, finely striated bosses or mamelons. The most caudad of these prominences lies 250u in front of the cloacal aperture and is 156 to 187» long; the second lies 140 to 155 anterior of the first and is 125 to 155u long; and the third is 95 to 125» anterior of the second and is 125 to 140y long.
Female 11 to 14 mm. long and with a maximum width of 357 to 430. The esophagus is 435y long by 45y wide at the anterior end and increases to 981 in width at the posterior end. The esophageal bulb is 117 to 310% wide. The nerve ring surrounds the esophagus about 120u from the anterior end of the body. The cephalic dilation is 120 to 170 wide in outside diameter. The cervical papillae are sit- uated 260 to 280% from the anterior end of the body. The excretory pore is situated about 470u caudad of the union of the intestine and esophageal bulb. The lateral alae begin at the cervical papillae and extend to the posterior end of the body. The anus is situated 3 to 4 mm. from the tip of the tail. The vulva is situated 1.6 mm. from the anterior end of the body. (Fig. 5.) In immature specimens the vulva is covered with a brownish colored plaque; in gravid specimens, however, the vagina is protruded. Eggs oval, flattened on one side, and 96 to 98u long by 28 to 38y wide.
Host.—Glaucomys volans volans.
Location—Cecum and colon.
Locality —F¥ alls Church, Va.
Type specimens—United States National Museum Helmintho- logical Collections No. 27827; paratypes No. 27793.
Syphacia thompsoni resembles S. obvelata (Rudolphi, 1802), the type of the genus, in body form and in the number of ventral mame-
ART. 4 NEW NEMATODE WORMS FROM RODEN TS—PRICE a
lons in the male. It differs, however, in that the former species is about twice the size of the latter, the tail of both sexes is proportion- ally longer and more slender, the spicule and gubernaculum larger, and the egg smaller. The presence of a spiny process caudad of the cloacal aperture also appears to be characteristic of the new species. This species is readily distinguishable from S. pallaryi (Seurat, 1915) by the number of ventral mamelons, there being but two in the latter species.
Yorke and Maplestone (1926) list S. bonnet Thiel, 1925, from Mycetes seniculus, and S. stossichi (Setti, 1897) from Hystrix cris- tata, in addition to S. obvelata and S. pallaryi, as belonging to the genus Syphacia. Travassos (1925), however, has shown that S. bonnei is identical with Hnterobius minutus (Schneider, 1866). The inclusion of S. stossichi (=Oxyuris stossichi) in this genus by Tra- vassos (1923) appears to the writer to be subject to question. ‘The bosses or mamelons figured by Setti (1897) are too small and sit- uated too near the cloacal aperture to be considered comparable to the large mamelons of the other species of the genus; there also ap- pears to be an absence of caudal alae, papillae, and a gubernaculum. On the basis of these differences it is the opinion of the writer that if S. stossichi is restudied, it will probably be found to represent a new genus.
HELIGMOSTRONGYLUS HASSALLI, new species
Specific diagnosis —Heligmostrongylus: Small slender worms, reddish in color and loosely coiled when fresh, yellowish and almost straight when preserved in alcohol. Cuticle of the anterior end dilated and coarsely striated transversely. (Fig. 6.) The cuticle of the body is finely striated transversely and is also provided with small striated bosses (fig. 7) arranged in the form of 14 to 16 longi- tudinal lines. These lines extend from the cephalic dilation to the vulva of the female, and to the bursa of the male. A well-defined ala extends along the left side of the body from the cephalic dilation to the vulva in the female, and to near the bursa in the male. Cervical papillae not observed. The mouth opening is triangular and bears 2 submedian papillae, and one amphid (the lateral papilla of various authors) on each side. (Fig. 8.)
Male 5.3 to 6.8 mm. long by a maximum width of 122» at the middle of the body. The cephalic expansion is 38. wide and 75p long. The esophagus is simple, 290 to 300» long by 15 wide at the anterior end and increasing to 40x in width at the posterior end. The nerve ring surrounds the esophagus about 150% from the an- terior end. The excretory pore is situated immediately caudad of the posterior end of the esophagus. The bursa consists of two large
4 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
lateral lobes and a smaller dorsal lobe. The ventro-ventral and latero- ventral rays are widely separated and divergent, the latero-ventral ray being longer than the ventro-ventral and extending to the edge of the bursa; externo-lateral ray slightly thicker than the other rays; medio-lateral ray relatively thick and extending almost to the edge of the bursa; externo-lateral ray short and divergent; externo- dorsal ray slender; dorsal ray cleft almost to the base, forming two branches which in turn give off a lateral branch near the origin of the primary branch. (Fig. 9.) Prebursal papille present. The spicules are tubular, pointed, and 385, long by 16m wide at the base. Gubernaculum absent. The telamon (fig. 10) is feebly chitinized and elongated; its anterior end pointed and embedded in the ventral wall of the cloaca; the lateral edges are inrolled, forming a tube through which the spicules pass; slightly caudad of the middle of the telamon a slender lateral process is given off which appears to furnish attachment for muscles.
Female 8.4 to 8.6 mm. long and 76 wide at the vulva. The esoph- agus is 350u long by 20» wide at the anterior end and the width increases to 50» at the widest portion. The nerve ring is situated 210» from the anterior end of the body. The tail is short and blunt. (Fig. 11.) The anus is situated about 53» from the tip of the tail. The uterus is single and is provided at its distal end with a strongly muscular ovejector which measures 220u in length. The vulva is situated about 180. from the posterior end of the body. In some specimens a short, thick, ventral process is present immediately in front of the vulva. Egg 76» long by 45, wide, oval in shape and provided with a thin shell.
Host.—Seciurus carolinensis.
Location—Small intestine.
Locality —Bowie, Maryland.
Type specimens—United States National Museum Helmintho- logical Collections No. 27853; paratypes No. 27860.
This sepcies differs from Heligmostrongylus sedecimradiaius (Linstow 1899) in its smaller size, shorter spicules and dorsal ray. The dorsal ray in H. hassalli is not cleft entirely to the base and the lateral branches arise near the bifurcation. In H. sedecimradiatus the dorsal ray is cleft entirely to its base forming a double dorsal ray, and the lateral branches are given off about the middle of the primary branches. Travassos (1921) says that a gubernaculum is present in S. sedecimradia‘us, but from his figure of the bursa of this species the shape and position of this structure suggest that it functions as a telamon and that a gubernaculum is absent.
ART. 4 NEW NEMATODE WORMS FROM RODENTS—PRICE 5
REFERENCES
SETTI, ERNESTO. 1897.—Nuovi elminti dell’ Hritrea. Boll. mus. di zool. (ete.), Genova (vol. 57), 50 pp., pls. 8-9, 41 figs. SeuraT, L. G. 1915.—Sur deux nouveaux oxyures du Maroc. Bull. Soc. d’hist. nat. de l’Afrique du nord, Alger, vol. 7 (2), 15 fev., pp. 24-81, figs. 1-9. TRAVASSOS, LAURO. 1921.—Contribuicdes para o conhecimento da fauna helmintolojica bra- sileira. XIII. Ensaio monografico do familia Trichostrongylidae Leiper, 1909. Mem. Inst. Oswaldo Cruz, Rio de Jan. Manguinhos, vol. 138 (1), pp. 1-135, pls. 1-56, figs. 1-250. 1923.—Informacds sobre a fauna helminthologica de Matto Grosso. Folha Med., Rio de Jan., vol. 4 (5), p. 35. 1925—Fauna brasiliense. Nematodes. Oayuroidea-Oxryuridae. Revisio do genero Hnterobius Leach, 1858. Mus. vac. do Rio de Jan., new ser., No. 2, 11 pp., 2 pls., figs. 1-15. YORKE, WARRINGTON; and MApLESTONE, P. A. -. 1926.—The nematode parasites of vertebrates. With a foreword by C. W. Stiles. xi+-536 pp., 307 figs. London.
EXPLANATION OF PLATES
a=anus. oesb=esophageal bulb. e€ pp=excretory pore. ovj=ovejector. int=intestine. ut=uterus. la=lateral ala. vag=vagina. nr=nerve ring. vul=vulva.
oes=esophagus.
PLATE 1 Syphacia thompsoni, new species
Fig. 1. Head, end face view.
. Adult male, entire.
. Male, posterior end.
. Spicule and gubernaculum.,
. Female, anterior end.
oR WON HE
PLATE 2 Heligmostrongylus hassalli, new species
Fic. 6. Female, anterior end. 7. Cuticular bosses; greatly enlarged. 8. Head, end face view. 9. Bursa, dorsal view. 10. Telamon, ventral view; greatly enlarged. 11. Female, posterior extremity.
O
ba GE i. og o sin
ae i | : ee fi (y $e ” he ; | ; , | _
U. S. NATIONAL MUSEUM PROCEEDINGS, VOL. 74, ART. 4. PL. 1
SYPHACIA THOMPSONI, NEW SPECIES
FOR EXPLANATION OF PLATE SEE PAGE 5
U. S. NATIONAL MUSEUM PROCEEDINGS, VOL. 74, ART. 4. PL. 2
LVOTHTA ‘
HELIGMOSTRONGYLUS HASSALLI, NEW SPECIES
Jom
FOR EXPLANATION OF PLATE SEE PAGE 5
FOSSIL FOOTPRINTS FROM THE FORT UNION (PALEOCENE) OF MONTANA
By Cuaries W. GILMorE
Curator of Vertebrate Paleontology, United States National Museum
INTRODUCTION
in 1908 Mr. A. C. Siberling, while collecting mammalian fossils
for the United States National Museum in the Fort Union deposits of south-central Montana, also discovered a series of fossil footprints in this same formation.
Apparently these were the first fossil tracks to be discovered in the Paleocene, and as such they appear worthy of record. Recently a series of fossil tracks found in deposits of equivalent age in the ‘Province of Alberta, Canada, have been described by Messrs. Ruther- ford and Russell,’ cee these tracks are thought to be mammalian in origin, and alata wets their details are somewhat obscurely preserved their tridactyle nature effectually distinguishes them from the tracks here considered.
Originally the Montana tracks were preserved as one large slab, but owing to the vicissitudes of transport this specimen was broken into many pieces and the loss of connecting edges made it impossible to reassemble them in their original relationships. The specimen is now in three slabs, as shown in the accompanying plates.
The tracks are impressed on the slightly undulating surface of a fine grained sandstone that in some instances has preserved the full details of the feet in addition to recording dragging tails, claw scratches, and belly impressions. Due to the breakage of the original slab, only short sections of trackways are now available. The best one at hand, shown in Plate i, is selected as the type.
AMMOBATRACHUS MONTANENSIS, new species
Plates 1, 2, and 3
Type.—Cat. No. 7635, U.S.N.M.; parts of trackways on three slabs that originally formed one large slab. Collected by A. C. Silberling, 1908.
Type locality—Section 8, range 16, township 5, Bear Butte, Bear Butte Pass, Sweetgrass County, Mont.
Geological horizon Silberling’s Fort Union No. 3.” Paleocene.
1 Rutherford, R. L. and Russell, L. S., Amer. Journ. Sci., vol. 15, 1928, p. 262.
No. 2750.—PRocEEDINGS U. S. NATIONAL Museum, VoL. 74 ART. 5 3970—28 1
2 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
The series of tracks selected as being typical of the above-named species is shown on the lower two thirds of the slab illustrated in Plate 1. With the exception of four faint impressions shown on the bottom of Plate 2, all other tracks on the three slabs illustrated in Plates 1, 2, and 3 are regarded as pertaining to the present genus and species.
Description.—Stride vari- able, measuring from 135 to 180 mm.; width of track- way measured from the out- side of one foot to a cor- responding position on the opposite side, about 162 mm. The manus tracks irregu- larly placed but usually for- ward of midway between those made by the hind feet. Fore slightly smaller than hind foot.
Manus.—Tetradactyle, plantigrade, rounded palm, and diverging digits. Length over all, 22 mm.; distance between tips of lat- eral toes 31 mm.; breadth of palm, 20 mm, Outer toe originates well backward on the side of the palm and is strongly diverted outward away from the three inner toes. The manus as a whole turns strongly inward toward the median line of the trackway. Inner toe shorter than fourth digit.
Fic. 1—AMMOBATRACHUS MONTANENSIS, TYPE No. Third longest. 5 SacN; i d ‘ 7635 U. N. M. DtaGraMm OF TRACK WAY Pes —Pentadactyle, plan-
ABOUT ONE-THIRD NATURAL SIzH
tigrade. Sole broadly rounded behind. Length over all, 33 mm.; distance between tips of lateral toes, 33.5 mm.; breadth of sole, about 28mm. _ First digit short and originating well back on the side of the sole; divercation slightly less than 45°. Three median toes long, relatively slender, with sub- acute or rounded terminations. There is some variation in the rela- itive lengths of the digits of opposite feet as is clearly indicated in Figure 1. None of the tracks give clear indication of the presence of
ART. 5 FOSSIL FOOTPRINTS FROM MONTANA—GILMORE a
sharp claws although deep scratches are recorded by the toes espe- cially of the pes where they dragged with each step. These are clearly indicated on the left side of the trackway shown in Plate 1. In this same section of trackway a smoothing out of the surface be- tween the two lines of tracks suggest a belly drag leading to the inference that the animal was a low, wide-bodied, short-legged quad- ruped. The presence of a heavy tail is clearly indicated by an in- termittent but deep median groove.
The digital formula of four and five toes, respectively, on manus and pes, a close similarity in arrangement and in the relative iengths of the digits, these tracks have their closest affinities with Ammobatrachus turbatans Gilmore recently described* from the Supai formation (?Pennsylvanian) of the Grand Canyon. ‘Their specific distinctness, however, is at once indicated by the much greater width of the trackway, longer stride, and more open spac- ‘ing of the fore and hind foot impressions. The larger foot measure- ments of A. turbatans suggests a bigger animal than the one mak- ing the tracks considered here, ‘which accentuates the importance of the differences enumerated.
Comparison of the pes tracks shows the Montana ichnite to have relatively longer and more slender digits with a more strongly divergent digit five. Contrasting the manus tracks the sole of Cat. No. 7635, U. S. N. M., is shorter, the lateral digits relatively longer, and the angulation of the imprint with toes directed strongly inward toward the center of the trackway, at once distinguishes these imprints from the forwardly pointed toes of A. turbatans.
The digital formula strongly suggests this new species to be of amphibian lineage, but in the absence of confirmatory evidence there seems no way of definitely determining this point. Neither does a review of the known fauna of this formation give any hint as to the class or the kind of animal to which these tracks might be attributed. There are no amphibians known from the Fort Union and the only reptiles are small lizards, turtles, crocodiles, and the Rhyncocephalian Champsosaurus. The two last mentioned with their elongated toes of the hind feet would be at once ruled out, though there is the possibility of their being Chelonian in origin.
2Smith. Misc. Coll., vol. 80, 1928, No. 2956, p. 8, pl. 2.
EXPLANATION OF PLATES
PLATE 1
Ammobatrachus montanensis, new species. No. 7635 U.S.N.M. type. Trackway on lower half of slab shows toe scratches, belly drag and tail grooves. About one-third natural size.
PLATE 2
Ammobatrachus montanensis, No. 7635 U.S.N.M. Part of type slab. Various imprints of feet. Tracks crossing the slab diagonally at the bottom pertain to some other animal. About one-third natural size.
PLATE 3
Ammobatrachus montanensis, No. 7635 U.S.N.M. Part of type slab. Various imprints of feet with distinct tail drag. About natural size.
4 C
Plead
PROCEEDINGS, VOL. 74, ART. 5,
U. S. NATIONAL MUSEUM
TYPE SLAB OF AMMOBATRACHUS MONTANENSIS
FOR DESCRIPTION OF PLATE SEE PAGE 4
PE
PROCEEDINGS, VOL. 74, ART. 5.
U. S. NATIONAL MUSEUM
PART OF TYPE SLAB OF AMMOBATRACHUS MONTANENSIS
FOR DESCRIPTION OF PLATE SEE PAGE 4
U. S, NATIONAL MUSEUM PROCEEDINGS, VOL. 74, ART. 5, PL. 3
PART OF TYPE SLAB OF AMMOBATRACHUS MONTANENSIS
FOR DESCRIPTION OF PLATE SEE PAGE 4
NOTES ON SOME ORIENTAL SAPROMYZID FLIES (DIP- TERA), WITH PARTICULAR REFERENCE TO THE PHILIPPINE SPECIES
By J. R. Matiocu
Of the Biological Survey, United States Department of Agriculture
For many years I have taken a great interest in the family Sapro- myzide, collecting the species assiduously in Scotland, where they occur in great numbers, and during the past decade I have devoted much time to a consideration of the generic groupings. Recently I published several papers on the American and Australian forms, and at my request Dr. J. C. H. de Meijere has submitted for exam- ination types or paratypes of most of his Oriental species,’ along with specimens of a number of other species from the same region described by different authors. This material, coupled with a large collection from Formosa sent to me by Dr. Walther Horn, has en- abled me to arrive at a definite conclusion as to the generic positions of these species and the specific identities of many Philippine species submitted to me for identification by the late Prof. C. F. Baker.
The work on the genera of the family by Dr. F. Hendel * appeared at a time when generic concepts in the group were much broader than they are to-day, a fact clearly shown by the acceptance then of but 22 valid genera, as compared with 76 in the same author’s generic synopsis published in 1925. Unfortunately the broad concepts laid down in 1908 have influenced most workers on the family so that species have been described in the genus Lauwania which have very little in common with the genotype. While I do not favor the erec- tion of a large number of monobasic or poorly represented genera in any group, I feel that a careful consideration of structural charac- ters by a competent taxonomist ought to result in a well ordered segregation of related forms in genera, or subgenera, the distribu- tion of which throughout the various faunal regions can not fail to be of interest and value in affording data upon the relationships and
1¥For list, see Tijds. Ento., vol. 60, 1918, p. 345. 2 Genera Insectorum, fase. 68, 1908.
No. 2751.—PRocEEDINGS U. S. NATIONAL Museum, VOL. 74, ART. 6. 2609-291 1
2 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
origin of these groups, and their connections with others found in the same faunal regions. In other words, I consider that a genus should contain closely related forms, how close being a matter for decision by a number of competent workers, and not a heterogenous collection of diverse forms, so that the occurrence of a species of such genus will have more significance than where a member of a heterogenous concept occurs.
The most comprehensive paper on Philippine Sapromyzidae is that by Dr. R. Frey which appeared in 1927.* It contains records of 54 species, 27 of them new to science. I have gone carefully over this paper and have succeeded in identifying a number of the species in the material in my possession, but several are yet unknown to me. In the present paper I indicate certain synonyms as the result of Doctor Frey’s work and also relocate some of his species. The genera unknown to me are briefly discussed also.
While all of the systematic entomological work on Philippine species has up to the present been done by European and American workers, it is certain that in the near future some capable Philippine students will undertake this work; and in the present paper I have attempted to utilize the most dependable characters for the separation of the species so that even without access to type specimens it may be possible for a careful and efficient worker to confidently identify those included in this paper.
The reason for introducing so many extralimital species in the key is that, though as yet unrecorded from the Philippines, it is not at all certain that many of them do not occur there, because the col- lecting that has been done has been by no means intensive, and the fact that about twice as many species are known from the Nederland Indies as are recorded from the Philippines appears to me a safe premise from which to deduce that many more Philippine species have yet to be discovered, and probably many of these will be iden- tical with species occurring in adjacent or even distant islands in the same region.
The figures of the male hypopigia are presented as the most depend- able specific indices, and this series is the most extensive ever pub- lished for this family.
The original intention was to publish this paper in the Philippine Journal of Science, but this plan was changed by the death of Prof. C. F. Baker and the acquisition of his collection by the United States National Museum. To make the collection as complete as pos- sible, I waive any claim to the type specimens of the Philippine species collected by Professor Baker and deposit them in the National Collection.
* Acta Soc. Faun. Flor. Fennica, vol. 56, No. 8, pp. 44.
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—-MALLOCH 3
Subfamily CELYPHINAB
This group has usually been given distinct family rank, but I con- sider it is merely a subfamily of Sapromyzidae, elise omrcnied by the very large convex scutellum, which usually covers the entire abdomen and gives the insects a beetle-like appearance, and is with one exception without marginal or discal bristles. With the discov- ery of the new genus /diocelyphus described herein the claim to family distinction of the group is very much weakened as in it the scutellum is very much smaller than usual and it has four well- developed bristles. Many of the species are metallic blue or violet colored, quite distinct from any in the other subfamily, but some are almost entirely testaceous, a color predominating in the Sapro- myzinae. I have recently briefly discussed the family characters in “Entomologische Mittelungen” (1927, page 160), but did not at that time have access to the new genus above mentioned. In addi- tion to the characters mentioned in that paper it appears worth noting that while there are no distinct bristles on the frontal field there are four very fine minute hairs which appear to me to repre- sent the two pairs of orbital bristles usually present in Sapromyzinae. One of these pairs of fine hairs is situated near the margin on upper half and represents the upper pair of orbitals, while the other is near anterior margin and each hair is about as far from eye as from each other and incurved. In this subfamily we find also the only case where the cross vein separating the discal and posterior basal cells of the wings is absent in Sapromyzidae, but it is not invariably so in the group, being confined to three species previously placed in Spaniocelyphus and the single species of Idiocelyphus. The presence or absence of this cross vein has been utilized as a generic character in related families and is generally considered as of considerable im- portance in classification so I have deemed it proper to separate the three species first above mentioned from the typical forms of Spanio- celyphus.
The five genera at present known to me may be distinguished as below.
KBY TO THE GENERA
1. A distinct cross vein separating the discal and posterior basal cells of
No cross vein separating discal and posterior basal cells of wings_________ 4,
2. Arista very Slightly widened at base, the widest portion not more than one- fourth as. wide as third antennal segment; vertex rounded; postvertical pristless/absent he 2 ries etal Seo Se ae eae Paracelyphus Bigot. Artista very conspicuously widened on more than its basal half, leaf-like, its greatest width almost, or quite, as great as that of third antennal seg-
4 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. T4
8. Vertex rounded, postvertical bristles absent or microscopic. Celyphus Dalmann. Vertex sharply carinate above and slightly raised behind ocelli; postvertical bristles usually represented by microscopic hairs__ Spaniocelyphus Hendel. 4, Secutellum much longer than thorax, and without bristles; mesonotum without central bristles and the margin with but two, which are confined to the postsutural SCCHLOT LEAs eh ee ee Se Acelyphus, new genus. Scutellum not longer than thorax, with two bristles on disk near base and two on margin near apex; mesonotum with several pairs of dorsocentral and one pair of acrostichal bristles in front of scutellum, the humeral and notopleural bristles also present____________ Idiocelyphus, new genus.
ACELYPHUS, new genus
Generic characters —Most closely related to Spaniocelyphus, dis- tinguished from it by the presence of a well-developed pair of post- vertical bristles, the lack of a cross vein separating the discal and posterior basal cells of wing, and the much broader abdomen, which has the tergites evenly rounded over sides and without a slight suture about at the part where they curve over. In Spaniocelyphus there is a rather evident suture on each side of each tergite which divide it into three almost equal portions.
Genotype—Acelyphus politus, new species.
There are but three species known to me, which may be distin- guished as below.
KBY TO THE SPECIES
1. Entire disk of scutellum and mesonotum with dense erect miscroscopic pile; length of scutellum not over 1.5 as long as its greatest width; hypopygium
AS HI SURES. Oras Gat net ae ee ee oe ee eee stigmaticus (Hendel). Scutellum without dense erect microscopic pile, its length distinctly over 1.5 asHongias its: greatest widths tse aes che ae 2 ee 2
2. Scutellum smooth except for the widely separated piliferous depressions ; hypopygium as in Figures 9 and 10; mesonotum without microscopic erect pile except on extreme anterior margin__-____-_____~_ politus, new species.
Scutellum with regular close stipplelike punctures in addition to the pilif- erous depressions; mesonotum with dense erect microscopic pile on entire SUE CO ae ek Sh eee 2 ha repletus, new species.
ACELYPHUS POLITUS, new species
Male and female——Prevailing color tawny yellow, deeply over- laid with metallic violet-blue. Legs tawny yellow. Apices of palpi deep black.
Frons about 1.5 as wide as long, the vertex not very sharply carinate, almost rounded except just behind the ocelli and near each eye, the two pairs of verticals and the postverticals well de- veloped; flattened part of the arista fully twice as long as the apical hairlike parts; palpi much dilated at apices, spoon shaped. Scutellum without pronounced punctures, with almost imperceptible
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—MALLOCH 5
shallow depressions at bases of the very fine sparse hairs which occur over the entire surface. Hypopygium as Figures 9 and 10. Inner cross vein of wing a little beyond middle of discal cell; marginal cell not as wide as submarginal at apex, the second vein not very abruptly bent forward at apex.
Length, 4 mm.
Type.—Male, allotype, and 14 paratypes, Mount Maquiling, Luzon (C. F. Baker).
Type.—Cat. No. 41084, U.S.N.M.
Two of the paratypes are immature and appear to the naked eye almost uniform fulvous in color, but there is a very evident violet- blue tinge on the scutellum when they are viewed from behind.
ACELYPHUS REPLETUS, new species
Male and female——A darker species than the preceding one, more purplish than violet blue, with the same colored palpi.
Head similar to politus. A very striking character apart from _the double punctuation of the scutellum, is the presence of dense erect miscroscopic pile on the mesonotum. Similar pile occurs on the mesonotum and entire scutellum of stigmaticus, but there is no indication of such pile on the scutellum, and only a vestige of it on the extreme anterior margin of mesonotum, in politus. Inner cross vein beyond middle of discal cell; second vein rather abruptly bent forward at apex.
Length, 4 mm.
Type—K¥emale, Wai Lima, Sumatra, 1912 (Karny and Siebers), in author’s collection; allotype, male, and three paratypes, Singapore, Straits Settlements (C. F. Baker), in United States National Museum.
Allotype.—Male, Cat. No. 41087, U.S.N.M.
There is a possibility, but no certainty, that some of the smaller species of this group described as belonging to Celyphus may be
referable here. ACELYPHUS STIGMATICUS (Hendel)
Spaniocelyphus stigmaticus HENDEL, Suppl. Ent., No. 3, 1914, p. 93.
This species has occurred only in Formosa up to the present, but it may be found in other sections of the Orient. It was described when the genus Spaniocelyphus was erected, but scutatus Wiedemann was selected as the genotype.
IDIOCELYPEUS, new genus
General characters——Head similar to that of Spaniocelyphus, the vertex rather sharp, with four bristles and a pair of small, but dis- tinct, postverticals; oscellars lacking; antennae as in Spaniocelyphus;
6 PROCEEDINGS OF THE NATIONAL MUSEUM | vou. T4
clypeus projecting more than in the other genera, angular in profile. Thorax with one pair of quite well developed prescutellar dorso- centrals, and from one to three much shorter pairs in front of them, becoming progressively shorter anteriorly; the other bristles consist of: 1 humeral, 2 notopleurals, 1 supra-alar, 2 postalars, one pair of prescutellar acrostichals, and one mesopleural; scutellum not longer than.thorax on dorsum, gradually widened from base to middle, from there slightly narrowed, and broadly rounded at apex; basal bristles situated on disk, separated by about one-third of the width of scu- tellum, and not over one-fourth from base, the apical pair situated on extreme edge about one-third from apex, and upwardly directed. Legs as in the other genera, but the hind tibia has a strong, black, slightly curved, apical ventral spur which is almost as long as hind metatarsus. Wings as in the other genera, the discal cell not sep- arated from basal cell by a cross vein. Genotype—The following species.
IDIOCCELYPHUS BAKERI, new species
Shining brownish testaceous; clypeus violet-blue in front; dorsum of head and thorax with a violet tinge, the abdomen more bluish; all bristles black. Palpi with their apices narrowly black. Wings yellowish hyaline. Halteres yellow.
Frons smooth, subquadrate, with a pair of fine convergent hairs close to middle of disk; basal segment of antennae a little longer than second, bare below, third slightly tapered apically, about 1.5 as long as first and second combined; basal wide portion of arista not as wide as third antennal segment and a little longer than it and also than apical hair-like portion; cheek higher than width of third an- tennal segment; palpi not dilated. Thorax smooth on dorsum, with fine black hairs on mesonotum; scutellum smooth, without hairs, but with dense microscopic pile which is present also on thorax. Fore femur with two or three posteroventral bristles.
Length, 3.5 mm.
Type and three paratypes, Mount Maquiling, Luzon, P. I. (C. F. Baker).
Named in honor of the collector.
Type.—Male, Cat. No. 41073, U.S.N.M.
This genus supplies a connecting link between the Celyphinae and Sapromyzinae, the small size of the scutellum, and the presence of scutellar bristles, showing an approach to some genera in the latter subfamily. No other species in Celyphinae has scutellar bristles.
Genus PARCELYPHUS Bigot
I have before me two species of this genus, one of which appears to be undescribed. They may be distinguished as below.
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—MALLOCH if
KEY TO THB SPHCIES
1. Metallic dark blue species, with decided violet tinge, legs black, bases of tarsi yellow; scutellum with a shallow furrow on each side at base in addition to the one along marginal rim, the surface elsewhere smooth; posterior notopleural bristle present_--_-----__- sumatrensis van der Wulp.
Testaceous yellow species, the legs concolorous, scutellum with an almost imperceptible bluish tinge, without furrow at base except the marginal one, its surface irregularly rugose, or coarsely wrinkled; posterior notopleural BristlevaDSente= = ae en ee testaceus, new species.
PARACELYPHUS SUMATRENSIS van der Wulp
Paracelyphus sumatrensis VAN DER Wotp, Compt. rend, Soc. Hnt. Belg., p. 297, 1884.
This species must be very similar to hyacinthus Bigot, differing mainly in the color of the antennae, which are entirely black as against the yellow color of those of hyacinthus, though the third segment in the latter is brownish or fuscous.
Length, 6 mm.
Locality, Wai Lima, Sumatra, 1921 (Karny and Siebers).
PARACELYPHUS TESTACEUS, new species
This species is about the same size and build as the preceding one, but is readily distinguished by the general testaceous color, even the antennae and legs being pale. While the pale metatarsi of sumatrensis are distinctly thickened they are not at all thickened in testaceus. Other characters as in the key.
Length, 6 mm, width 4 mm.
Type, Mount Maquiling, Luzon (C. F. Baker); paratypes, one, Manila, (G. Compere); four, Tangcolan, Bukidnon; two, Butuan, Mindanao; one Mount Banahao; one, Davao, Mindanao; one, Samar Island; all in the Philippines (C. F. Baker).
Type.—Cat. No. 41087, U.S.N.M.
The first mentioned paratype has a label on it bearing the nota- tion ‘This is always to"be noticed round orange or lemon trees. Compere.’
Genus CELYPHUS Dalmann
Until recently all the species of the subfamily except those assigned to the preceding genus were located in Celyphus, but a few years ago Hendel proposed the removal of certain species to the new genus Spaniocelyphus, and this division is adopted herein as indicated in the generic key. It is not possible, however, to definitely decide the generic position of some of the previously described species. Those I now have before me, or can definitely locate, are included
8 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
in the keys given herein. All the material in the Baker collection, now a part of the collections of the United States National Museum, has been used in connection with this paper.
KEY TO THE SPECIES
1. Frons with only the outer, outwardly curved, vertical bristles present, and these very weak; thickened part of arista about 1.5 as long as third an- tennal segment, the apical hairlike portion about one-third as long as the wide portion; head and thorax fulvous yellow, scutellum violet-blue, slightly wrinkled-onsside@s =.= ==>. 's-:2_ 2 eee bisetosus, new species.
Frons with four quite strong and distinct vertical bristles; the wide part of arista but little longer than third antennal segment, the apical hair-like part fully half as long as the wide part; scutellum distinctly wrinkled
on?-most,OL Its suntacen eon eee ee ee ee ee ee ee 2. 2. Face with a black mark between each antenna and eye____-____________ oe Face without a black mark between each antenna and eye______________ -t
3. Both thorax and abdomen fulvous yellow, with distinct violet tinge; apical portion of arista not as long as the wide basal part, the latter not as wide as third antennal segment 2)» 2\) Sse. eis Ee puncticeps, new species.
Thorax fulvous yellow, with slight violet or blue tinge, the scutellum entirely metallic blue; apical part of arista at least as long as the wide basal part,
the latter as wide as third antennal segment____________ aurora Karsch. 4. Species entirely testaceous in color, no conspicuous blue tinge present; cheek with a blue mark below eye_____________________ difficilis Malloch.
Thorax fulvous, with a distinct blue tinge, scutellum metallic blue. obtectus Dalmann.
CELYPHUS PUNCTICEPS, new species
Male and female.—Testaceous yellow, with a very pronounced vio- let-blue tinge on entire dorsum; the face with a deep black spot between each antenna and eye.
Structurally similar to odtectus, but the arista narrower at base. Scutellum irregularly punctured basally as in obtectus.
Length, 5 mm.
Type, male, allotype, and two paratypes, Singapore, Straits Settle- ments; paratypes, one, Penang; two, Porto Princess, Palawan (C. F. Baker.
Type.—Cat. No. 41083, U.S.N.M.
This species appears to be close to karschi Bigot, but without an examination of the type specimens it is impossible to be certain of most of the species described by the older authors.
_CELYPHUS BISETOSUS, new species
Female—Head and thorax shining fulvous yellow, with a faint bluish tinge; scutellum dark metallic violet-blue; abdomen dark castaneous above, yellow below. Antennae fulvous yellow; palpi yellow at bases, deep black on the apical broadened portions. Legs fulvous yellow, femora darker, fore pair brown. Wings yellowish hyaline. Halteres dull yellow.
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—MALLOCH 9
Vertex with a small outwardly curved bristle close to upper angle of each eye; antennae about as long as width of frons, basal segment slender, and about twice as long as second, third a little longer than first, as wide as, and about two-thirds as long as, the broad part of arista, the hairlike portion of latter but little longer than second antennal segment; cheek about half as high as eye; palpi much broadened, leaflike, at apices. Thorax smooth. Scutellum wrinkled at base. Legs normal.
Length, 5.75 mm.
Type.—Cotschin, India. (In author’s collection.)
CELYPHUS DIFFICILIS Malloch Celyphus difficilis MALLocH, Ent. Mitt., vol. 16, p. 161, 1927.
This species was described in the paper of mine already referred to herein, and is known only from Formosa.
CELYPHUS AURORA Karsch
Celyphus curora Karscou, Berlin, Ent. Zeitschr., vol. 28, p. 1738, 1883.
This species is brownish testaceous, with a conspicuous metallic blue sheen on face and frons, all of dorsum of thorax and scutellum, most pronounced on latter. The most noticeable character for dis- tinguishing the species is the deep black mark on each side of face between antenna and eye. The arista is as broad on basal portion as width of third antennal segment and the apical hairlike portion is as long as it and quite long haired. The frons in the specimen be- fore me is not depressed, but almost even and rounded on vertical margin, and the four vertical bristles are well developed. The abdomen and fore femora are testaceous. The scutellum is wrinkled on basal half only.
Length, 5.5 mm.
Locality, Langkat, Sumatra (Deut. Ent. Inst.).
CELYPHUS OBTECTUS Dalmann
Celyphus obtectus DALMANN, Anal. Ent., p. 32, 1823.
This species is very similar to the preceding one, but the head and thorax are more pronouncedly yellow testaceous and less tinged with blue, the scutellum is deeper blue and there are no dark spots on face. The frons is slightly depressed at vertex and the arista is different in structure, as stated in the key.
Length, 4.5 mm.
Locality, Ceylon (Dr. W. Horn).
10 PROCEEDINGS OF THE NATIONAL MUSEUM Vou. 74 Genus SPANIOCELYPHUS Hendel
This genus is limited in this paper to those species which have a sharp vertex without well-developed postvertical bristles, and the discal and posterior basal cells of wing separated by a distinct cross vein. In addition to these characters all species available to me have the palpi much less dilated at apices than have the species of Acely- phus, and there is a pseudosuture on each side of each abdominal ter- gite which divides the surface into three almost equal portions and causes them to bend rather sharply over, almost angularly so, in sharp contradistinction to those of Acelyphus. The frons is also shorter than it is in Acelyphus.
There appear to be five species in the material before me only three of which I consider are already described, though it is possible some of the others are among those so imperfectly described by some of the older authors that they can not be reliably identified.
Genotype—Celyphus scutatus Wiedemann.
KPY TO THE SPECIES
1. Scutellum shallowly and coarsely pitted on almost the entire surface, only the the APEX SMOOCHEL? f Lhs Lee _ eRe Ne eed” SERRE We AP ENA See, SRAM RAE RP 2, Scutellum entirely smooth or with but faint traces of shallow pitting basally 2s ne nt a il a Se ee ee Sl _ oye Bi vera ie sane ee 3. 2. Apical section of superior foreceps of male hypopygium long, tapering to tip (figs. 1 and 2); Philippine species__________.___ scutatus Wiedemann. Apical section of superior foreceps of male hypopygium short and broad, somewhat boot shaped (figs. 2 and 3) ; Formosan species. formosanus Malloch. 8. Anterior margin of thorax, including the humeri and propleura, fulvous; hypopygium of male as Figures 4 and 5________ sumatranus, new species. Thorax entirely metallic blue, not fulyous anteriorly except sometimes on the SUITS ODL care ak ea en 4. 4. Face and frons entirely deep metallic blue______ nigrocoeruleus, new species. Head fulvous, with blue tinge on face and frons______ laevis van der Wulp?
SPANIOCELYPHUS SCUTATUS Wiedemann Spaniocelyphus scutatus WIEDEMANN Aussereur. Zweifl. Ins., vol. 2, p. 601, 1830.
I have before me many specimens of this species from the Philip- pines, and present figures of the male hypopygium of one of these specimens. (Figs. 1, 2.)
Localities, Manila; Mount Maquiling, Luzon; Davao, Mindanao; and Tangcolan, Bukidnon (C. F. Baker) ; Rangoon, Burma, Marcli, 1927 (F. J. Meggitt.)
SPANIOCELYPHUS FORMOSANUS Malloch Spaniocelyphus formosanus Matiocn, Ent. Mitt. vol. 16, p. 161, 1927. This Formosan species was described in my paper already referred
to herein. I present now figures of the hypopygium of the male. (Figs. 3, 4.)
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—MALLOCH 11 SPANIOCELYPHUS SUMATRANUS, new species
Male.—Head shining fulvous yellow, with a slight purplish tinge on face and frons, no dark marks on face or cheeks; third antennal segment darkened above; palpi slightly darkened at apices. Thorax metallic blue, dorsum deep blue, humeral angles and extreme anterior margin fulvous; scutellum metallic blue, deep purple from base to beyond middle on disk, the apex blue. Abdomen black, with an aeneous or purplish tinge. Legs dull fulvous or tawny yellow, the mid and hind femora infuscated. Wings yellowish hyaline, quite noticeably yellow at apices.
Vertical margin quite sharp, ocelli in carina; the type has no verti- cal bristles remaining, but in the paratype there is a weak inner verti- cal on each side; face concave below middle in profile; labrum quite broadly exposed; antennae distinctly shorter than width of frons, basal segment hardly longer than second, third about as long as basal two combined; flat part of arista fully as long as third seg- ment and about as long as the apical hairlike portion. Scutellum smooth except for a few widely separated shallow piliferous punc- tures. Inner cross vein of wing close to middle of discal cell; marginal cell as wide as submarginal at apex, the second vein almost rectangularly bent forward atapex. Hypopygiumas Figures 7 and 8.
Length, 4 mm.
Type, Wai Lima, Sumatra, 1921 (Karny and Siebers) ; paratype, Kepahiang, Sumatra, 1,960 feet, November-December, 1925 (H. C. Kellers.)
Type.—In author’s collection.
Paratype.—Cat. No. 41085, U.S.N.M.
SPANIOCELYPHUS LAEVIS (van der Wulp)
Oelyphus laevis VAN DER WUuLP, Tijdschr. v. Ent., vol. 22, p. 53, 1881. Specimens which appear to belong to this species are very similar to sceutatus, differing essentially only in having the mesonotum and scutellum less punctate, the latter being evidently punctate only at base laterally. Localities, one specimen, Los Banos, 13 specimens, Mount Maquil- ing, Luzon (C. F. Baker).
SPANIOCELYPHUS NIGROCOERULEUS, new species
Female.—Head glossy black, with a blue tinge, frons at anterior lateral angles, cheeks, and lower part of occiput, brownish yellow, upper occiput dark brown; antennae, aristae, and palpi, fuscous. Thorax and scutellum glossy black, with a blue tinge, abdomen con- colorous. Legs brownish yellow, coxae and mid and hind femora
1Z PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
fuscous, tibiae triannulate with fuscous, most evident on hind pair. Wings grayish hyaline. Knobs of halteres fuscous.
Frons about twice as wide as long, raised behind and in front centrally, all vertical bristles present but small; facial depression a little below middle; arista with the basal portion about twice as long as apical. Thorax and scutellum absolutely smooth, each with some scattered microscopic hairs. Fifth abdominal sternite about twice as wide as long. Wings normal.
Length, 4.5 mm.
Type—Cat. No. 41086, U.S.N.M. Cuernos Mountains, Negros (C. F. Baker).
This is the darkest colored species now known to me. If nigrifacies de Meijere belongs to this genus it may be distinguished from it by the much smaller size, 2.5 mm. in length, and the dark steel-blue color of the frons.
Subfamily SAPROMYZINAE
Dr. F. Hendel has recently published a key to the genera of the entire family exclusive of the Celyphinae,* and Dr. R. Frey has in addition to this published a generic key to the species known to him® so that it appears almost unnecessary to give a generic key in this paper. There are however some differences of opinion as to the validity of certain of the genera and some elucidation is required to make matters clear to anyone intending to make a more intensive study of the Philippine or Oriental fauna.
Doctor Frey erected 6 new genera in his paper and so far as pos- sible I have dealt with these in the text following. Certain of Doctor Hendel’s genera I can not accept, and one or two others are given a different interpretation so that some notes at least are required to explain these points.
Genus STEGANOPSIS de Meijere
This genus in the strict sense is confined to the Eastern Hemisphere. There is a closely related form in the Americas and to make clear the distinctions between the two segregates I include the latter in the key given below. I incline to the opinion that Steganolauxania is entitled to no more than subgeneric distinction. All species of the genus known to me are included in the key.
KEY TO THE SPECIES
1. Anterior pair of orbital bristles directed backward; frons uniformly shining; face smooth and glossy, convex above middle (America).
Subgenus Steganolauxania Frey, latipennis Coquillett.
Anterior pair of orbital bristles directed inward; frons not uniformly
Bren ss. SELES) SAR LIL PERS SOE DS _v CAIAR eee eee ee ee 2.
4Encycl. Entomol., Diptera, vol. 2, fase. 3, 1925. ®* Acta Soc. Faun, Flor. Fennica. vol. 56, No. 8, 1927.
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—-MALLOCH 13
2. Longest hairs on arista not as long as width of third antennal segment; thorax with two series of intradorsocentral hairs; palpi black at apices; wings browned, more noticeably so along costa, their apices usually nar- rowly whitish from tip of second to tip of fourth vein; frons dull yellow, with a large velvety black ocellar mark, and brownish marks at bases of the orbital bristles; face usually with a large violet-black spot on each
SLO eager Osa cy CLC yy eae eee SD ee 3. Longest hairs on arista more than twice as long as width of third antennal SESMEM Ge eee SE ee ES Sea Sg VS Se eee. 5.
8. Wing tip not whitish hyaline; face unspotted; second wing vein not very GlOSCRLOACOSEAS = hess ete ee 3 Le ee divergens Frey.
Wing tip whitish hyaline; face spotted; second wing vein close to costa__ 4. 4. Abdomen largely glossy blackish; fore tibia at middle, and hind femora basally, more or less darkened; hypopygium as Figure 11. melanogaster (Thomson). Abdomen normally glossy testaceous yellow; fore tibia and hind femora not darkened; hypopygium as figure 12__________________ convergens Hendel. 5. Wings entirely yellowish hyaline; head yellow, marked as in the two next preceding species, but the palpi entirely yellow; legs yellow, a small dark preapical ventral mark on fore femur, and the apical four segments of fore tarsi dark; face smooth and glossy, evenly convex ; intradorsocentral hairs DES CTe AC eee eset cee eee wears eee eee tm ee minor (de Meijere). Wings conspicuously infuscated; thorax and abdomen black, pleura with, or without, a large yellow mark above; legs more extensively black; palpi and halteres fuscous; intradorsocentral hairs quadriseriate__________ 6. 6. Head largely yellow, face entirely so; second wing vein not closer to costa GHA eCORtHIT Gye imate eee See ee es ee Seal divergens Frey. Head largely or entirely black, face not entirely yellow; second wing vein Nearer cOuCOSsiAa, thankto whind vel = lake see ee ee a ae 7. Thoracic dorsum and pleura black, the former with series of spots and vittae, and the pleura with spots and streaks, of black color on a gray- dusted ground; face not rugose; wings fuscous, with irregular hyaline markings on disk, a rather large round hyaline spot beyond middle of first posterior cell, and a small one below it, and the narrow apices most conspicuous; inner cross vein about one-third from apex of discal cell (ACUTE rest ey) pe a a ee ee Be annulipes Malloch. Thoracic dorsum black, with or without gray dusted vittae, but without black dots; wings not marked as above, the inner cross vein close to middle
OTC CANNY Ce Lee ee er a certs er a et ah ne eee ot SU 8. Sehoreslersrentirely; black same oe eens ae le Te ies einer ee ea 9. Horevegsnotcentirelyablacks24 Patan ascend en are Nae ey ES 8 10.
9. Mesopleura yellow; apices of wings not hyaline______________ bakeri Bezzi. Mesopleura black; apices of wings hyaline________________ aterrima Frey.
10. Frons opaque yellow, the ocellar spot and a spot on each side at upper eye margin opaque black; tip of wing not hyaline; mid and hind tibiae yellow. pusilla Frey. Mrons: black,;. other characters) not as above.224 229 4) 11. 11. Center of frons entirely glossy black, ocellar spot and a large mark at an- terior margin on each side velvety black; apex of wing hyaline; mid and
hind tibiae pale, with black annuli; pleura and fore metatarsus black. buruensis Malloch. Center of frons not entirely glossy; pleura on upper half, and the fore meta- CENTS RV CILG Wmeteniees ene ee meet tr Meee ee ORNs Mae SE rie RG oe 2}
14 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
12. Thoracic dorsum unicolorous black, sometimes faintly gray vittate an- Kerio rly. s2t)_ Sch) 6 ae ihe aE ie et Pee See ee ee pupicola de Meijere.
Thoracie dorsum with 10 or 12 complete narrow pale gray-dusted vittae. multilineata de Meijere.
STEGANOPSIS CONVERGENS Hendel
Steganopsis convergens HENDEL, Suppl. Ent., vol. 2, p. 102, 1913.
I suspected that this species might be the same as melanogaster (Thomson), but the male hypopygia are entirely distinct as shown in Figures 11 and 12. The only other distinctions are those listed in the foregoing key.
I have not seen melanogaster from outside of Australia, but con- vergens occurs in Formosa and the Philippines. I have a male from Cuernos Mountains, Negros (C. F. Baker).
It appears evident to me that Pachycerina apicalis Bezzi is this species. Frey considers that the first name for the species is cwr- vinervis (Thomson).
STEGANOPSIS MINOR (de Meijere) Steganopsis minor DE MEJERE, Tijdschr. v. Ent., vol. 57, p. 237, 1914 (Pachycerina).
I have examined a specimen of this species sent to me by Doctor de Meijere and find that it belongs to this genus and not to Pachy- cerina in which it was described, and in which Frey retained it. It is known to me from Java, but Frey records it from Banahao, Luzon.
STEGANOPSIS BAKERI Bezzi
Steganopsis bakeri Brzzi, Philippine Journ. Sci., vol. 8, p. 315, 1913.
This species is unknown to me. Recorded only from Los Banos,
Paks STEGANOPSIS DIVERGENS Frey
Steganopsis divergens Frey, Acta Soc. pro Fauna et Flora Fennica, vol. 56, No. 8, p. 10, 1927.
As I have not seen this species and am uncertain of some of the characters of it I have inserted it in two places in the key. Described from Mount Banahao, Luzon.
STEGANOPSIS PUSILLA Frey
Steganopsis pusilla Frey, Acta Soe. pro Fauna et Flora Fennica, vol. 56, No. 8, p. 13, 1927.
Described from Limay, Luzon.
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—MALLOCH te
STEGANOPSIS MULTILINEATA de Meijere Steganopsis multilineata pr MertseRE, Tijdschr. v. Ent., vol. 67, 1924, p. 53.
I have examined the type specimen of this species sent to me by Doctor de Meijere and consider wndecimlineata Frey is synonymous with it.
Frey described his species from’ Banahao, Luzon, and Kolambugan, Leite.
This species and pupicola occur also in Ceylon (W. Horn).
Genus LYPEROMYIA Frey
This genus is unknown to me. It is closely allied to Steganopsis, being distinguished mainly by the presence of four equally strong pairs of dorsocentral bristles, one sternopleural, shorter second wing vein, and narrower frontal triangle.
Genotype.—Lyperomyia calopus Frey.
LYPEROMYIA CALOPUS Frey
_ Lyperomyia calopus Frey, Acta Soc. pro Fauna et Flora Fennica, vol. 56, No. 8, p. 15, 1927.
A black species, with gray-dusted spot on each orbit near antennae, yellow third antennal segment, and black fore legs, the latter with coxae, and a broad ring at middle of femora, yellow, apical third of tibia white, mid and hind legs yellow, coxae entirely, mid femora with exception of apices, hind femora entirely, and apices of tarsi, black. Wings yellow, bases brown, veins somewhat brownish clouded.
Described from Mount Polis, Luzon.
Genus XANGELINA Walker
This genus has a peculiarly shaped head (Figs. 13 and 14), the frons being very steep, the face broad and evenly, though slightly, convex and glossy, and the divisions below the eye exceptionally distinct. The anterior orbitals in basiguttata are almost indistin- guishable microscopic hairs, and are incurved, while the ocellars are lacking. The thorax has four pairs of dorsocentral bristles and acrostichals, the anterior pair of each being in front of the suture; there are two distinct sternopleurals; and the wing is as in Sapro- myza Fallen. There is no anteroventral comb on the fore femur, and all the tibiae have the preapical dorsal bristle distinct, the one on hind pair being long and fine.
XANGELINA BASIGUTTATA Walker
Xangelina basiguttata WALKER, Proc. Linn. Soe. London, vol. 1, p. 32, 1857.
A shining testaceous yellow species, with two black spots on apex of scutellum, and a black mark at apex of auxiliary vein of wing; inner cross vein of wing faintly clouded.
Length, 4.5 mm.
16 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
I have seen the species from Java, submitted by Doctor de Meijere.
Hendel does not include this genus in his key, in which it runs to Sciasmomyia Hendel, though evidently distinct from it, the latter having two pairs of equally long backwardly bent orbitals and a pair of strong ocellars, in addition to other differentiating characters. The African species Lauxania submetallica Loew belongs to A angelina.
PLEURIGONA, new genus
Generic characters —This genus will run down in Hendel’s key to genera to [chthyomyia de Meijere, but it is readily distinguished therefrom by the small conical projection of the propleura directly over the fore coxa, the lack of a propleural bristle, the much less prominent face, biemarginate vertical margin, lack of ocellar bristles, and the pronounced downward curvature of the apical section of the fourth wing vein. This last character distinguishes it also from any other genus of the family known to me at this time.
Genotype—The following species.
PLEURIGONA CURVINERVIS, new species
Male—Shining testaceous yellow, the type specimen greasy but with evidences of two dark submedian lines on anterior half of meso- notum. Wings quite noticeably yellow, veins brown, apices of first, second, and third, darker, but without a surrounding cloud.
Frons about 1.5 as wide as long in center, the vertex sharp and with a shallow emargination on each side of the ocellar triangle, posterior ocelli on edge of vertex, the four vertical and two post- vertical bristles quite well developed, ocellars lacking, both pairs of orbitals very weak and short, upper pair at almost their own length from eyes, the shorter anterior pair even farther from eyes; orbits not differentiated; surface of frons bare; face broad and evenly convex, shining, about 1.75 as high as frons; cheek about two- thirds as high as eye, the hairs fine and rather long; basal antennal segment very short, third about twice as long as wide; arista with sparse hairs, the longest about as long as width of third antennal segment; palpi slender. Thorax seen from the side pronouncedly arched, with 1 or 2+3 or 4 pairs of dorsocentrals and 2+4 pairs of acrostichals; the presutural, notopleural, and humeral bristles, quite fine, no sternopleural visible in type; scutellum pronouncedly convex, with four fine bristles. Abdomen short, the hypopygium small. Fore femur without an anteroventral comb; fore and hind tibiae with the preapical dorsal bristle quite long and fine, the mid tibia with that bristle short and strong. Inner cross vein at middle of discal cell; outer cross vein close to middle of wing; marginal cell rather a
5) and uniform, the second vein roundly curved forward at apex; third
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—MALLOCH 17
vein straight; fourth very conspicuously curved down from near middle of its apical section, the first posterior cell fully twice as wide at apex as at outer cross vein; costa as in Sapromyza.
Length, 5 mm.
Type.—Cat. No. 41075, U.S.N.M. Mount Maquiling, Luzon (C. F. Baker).
The head in this genus is very similar to that of Xangelina Walker, but in the latter the frons is not so sharp, nor is it biemarginate, and the upper orbital is long and strong, while the stigmatal region is also normal, with a strong bristle, and the fourth vein is but slightly divergent from third apically.
Genus AMPHICYPHUS de Meijere
This monobasic genus is readily distinguished from any other in the subfamily by the presence of an unequai pair of strong curved black spurs at apex of hind tibia on its anteroventral side, the longest of which is fully half as long as the basal segment of hind tarsus. ‘The hairs on the entire insect including those on the legs are very long and almost bristle-like, the third antennal segment tapers to the apex, the arista is short haired, the anterior orbital bristles are incurved, the scutellum is slightly tumid at bases of the apical bristles,‘and the costa has only fine hairs, no short black setu- lae being evident.
AMPHICYPHUS RETICULATUS (Doleschal)
Ensina reticulatus DoLEsScHAL, Natur. Tijdschr v. Nederl. Indie., p. 412, 1856.
Locality, Mount Maquiling, Luzon (C. F. Baker).
This species has been recorded from the Dutch East Indies and Australia, but this is the first record of its occurrence in the Philip- pines,
EUPROSOPOMYIA, new genus
Generic characters—Head in profile as in Figure 15, the face shining, and with a subconical production, below the level of which there is a transverse impression. The frons is similar to that of Prosopomyia Loew, having both the pairs of orbitals bent back- ward, and the surface with rather dense short hairs. In the type the ocellars are bent backward, but this may be due to injury. From Prosopomyia, to which genus it runs in Hendel’s recent key, the genus may be distinguished by the presence of several strong bristles close to the lower hind margin of each eye. These bristles are not on a callosity, which is the case in Awustralina Malloch, and the latter genus has the frons more than twice as long as wide and
2609—29—2
18 PROCEEDINGS OF THE SATIONAL MUSEUM VOL. T4
sparsely haired. The apices of the wings in the type specimen are damaged, but I believe the costa is the same as in Homoneura. Genotype-——The following species.
EUPROSOPOMYIA MACULOSA, new species
Male.—Head testaceous, frons opaque, with ocellar spot brown, and two brown submedian stripes which curve round the anterior extremities of the orbits and extend between those and the eyes; face glossy, with 9 black spots. One between and above the an- tennae, one between each antenna and eye, and a larger one between and below antennae, all velvety, a large glossy one on lower central part of convexity, and two on each side of it, the lower one not so glossy as the upper; a brown streak along the transverse impression ; antennae and palpi testaceous yellow. Ground color of thorax testaceous, largely obscured by fuscous markings, the dorsum with six subcontiguous vittae and the lateral margins fuscous, humeri with a central dark mark; pleurae largely fuscous; scutellum dark on each side and in center. Wings fuscous, with a pale mark on each side of inner cross vein, and the hind margin narrowly hyaline, the hyaline part bidentate in second posterior cell, and very narrow round apex of wing. MHalteres yellow.
Frons about 1.5 as long as wide, slightly widened anteriorly; arista sparsely short haired. Thorax with three pairs of postsutural dorso- centrals and about six series of intradorsocentral hairs, the median two series strong posteriorly; scutellum subconvex; both sternopleu- rals strong. Abdomen stout; hypopygium as Figure 16. Legs stout; fore femur with an anteroventral comb and short, rather irregular posteroventral bristles; mid tibia with one long and one short apical ventral bristle; hind femur with some preapical anteroventral setu- lae; fore and hind tibiae with short preapical dorsal bristle. Inner cross vein a little beyond middle of discal cell; penultimate section of fourth vein about four-fifths as long as ultimate section.
Length, 4 mm.
Type—Cat. No. 41601, U.S.N.M. Cuernos Mountains, Negros (C. F. Baker).
Genus PROSOPOPHORELLA de Meijere
This genus is readily distinguished by the shape of the head, which when seen from in front is as Figure 17, the process on middle of lower margin of ‘face, and the angular production of the cheeks being unique in this family. Another outstanding character is the presence of short hairs on the disk of the scutellum. Both pairs of orbitals are bent backward, and, though the face is noticeably convex, it is not at all shining but entirely dull. The ocellars are long, postver-
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—-MALLOCH 19
ticals short, and the arista short haired. Thorax with three pairs
of postsutural dorsocentral bristles and one sternopleural. Legs
rather long, fore and mid femora very much attenuated apically, the
former with an anteroventral comb, fore and mid tibiae and tarsi
slender, the preapical dorsal bristle distinct only on the hind tibiae. The genus is monobasic.
PROSOPOPHORELLA BUCCATA (de Meijere)
Prosopophora buccata pe Mertsrrn, Tijdschr. vy. Ent., vol. 53, p, 144, 1910.
Male.—Head testaceous. Thorax black, shining, pleura brownish and with some patches of whitish dust. Abdomen shining black. Legs testaceous. Wings ‘fuscous, with the following hyaline marks: An almost complete fascia proximad of the inner cross vein, a short fascia from costa to fourth vein proximad of outer cross vein, a quadrate mark at middle of first posterior cell, and another just proximad of it in second posterior cell, a mark over outer cross vein, and one at wing tip.
Length, 4.5 mm.
Habitat, Nederland Indies.
Genus PACHYCERINA Macquart
The characters cited by Hendel in his recently published key to the genera for the separation of this genus from Physogenta Mac- quart consist of the position of the anterior pair of orbitals, more closely placed than posterior pair in Physogenia and equally widely separated in Pachycerina, the differently haired aristae, shortly and densely haired in Pachycerina and longer and more sparsely haired in Physogenia, with slight differences in the form of face and occiput. Only the first mentioned character appears to be of any use for dis- tinguishing the genera as represented in the species before me. The only species of Physogenia which I have available now is ferruginea Schiner, and in the male of it the mid tarsus has the basal segment thickened, which is not the case in any of the species of Pachycerina available to me now. In my paper on Sumatran Sapromyzidae, which has already been referred to, I separated leucochaeta de Meijere from the other species previously included in this genus, erecting the genus Melanopachycerina for it and two other species. Doctor Frey retained this species in Pachycerina, but he also retained Steganopsis minor (de Meijere) in the genus so that he did not ap- parently have a very clear concept of the generic characters.
I can not determine to what genus Pachycerina cyaneostoma Frey belongs, but am doubtful if it belongs here. The following key will
serve to distinguish the species of the genus now known to me from the Orient.
20 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74 KEY TO THE SPECIES
1. Hairs on arista rather dense and not erect, the longest not as long as basal
width, :of) third: -antennaliseements: a 26 45225 s2¢_ feed el ee 2: Hairs on arista not dense, erect, and the longest very distinctly longer than basal width, of thirdsantennal .segmentie.. = ss = a eee 5.
2. Face without dark discal spots; a deep black velvety spot between each an- tenna and eye; third antennal segment shorter than usual, not over twice as long as its basal width; seventh abdominal tergite with a pair of black SUDO TS Sa a Be re ee oe ae ge ene ee sigillata (de Meijere).
Face with a pair of dark discal spots above middle; no black spot between each antenna and eye; third antennal segment elongate, about four times as‘long fas’ its! basal ‘widths 2S ee ee a ee eee 3.
38. Thorax with two series’ of intradorsocentral hairs, and four or five fine brownish lines on dorsum; fore femur entirely yellow; abdomen with a pair of small round black spots on seventh tergite______ javana Macquart.
Thorax with four series of intradorsocentral hairs, and ten fine black lines on dorsum; fore femur with a black apical mark; abdomen without a pair of black spots"on sevenuhotercites.. 22 2s ees a nee 4,
4. Abdomen broadly deep black on dorsum, narrowly yellow on sides. decemlineata de Meijere. Abdomen entirely ;yellow=s=-= 23-3 = eee flaviventris, new species.
5. Facial spots very small and pale brown in color; wings quite intensely yel- NO Wer BLOM ECON Gea ee ae ee ee sexlineata de Meijere
Facial spots larger and deep black; wings not more noticeably yellow along eostal than: elsewheres 22222. earn eee parvipunctata de Meijere.
PACHYCERINA JAVANA Macquart
Pachycerina javana Macquart Dipt. Exot., Suppl. 4, p. 274, 1850 (Sa- promyzda).
The largest species known to occur in the Orient, readily distin- guished from its allies by the characters listed in the key. The male has a pair of very long basal hypogygial forceps which taper apically, and extend forward below abdomen to well in front of its middle.
I have before me a series of specimens from Baguio, Benquet Province (C. F. Baker).
PACHYCERINA FLAVIVENTRIS, new species
This species is very similar to decemlineata, being the same in color except in having abdomen entirely yellow. The details in the above key are sufficient to insure its recognition.
The outer partial black vitta behind suture of thorax, as well as the two black pleural vittae, so distinct in decemlineata, are not visible in the type of flaviventris, and the pair of black spots on face are smaller than in that species.
T'ype—Cat. No. 41697, U.S.N.M.; Mount Maquiling, Luzon Proy- ince (C. F. Baker).
The other species listed in key I have seen only from Java; the species sigillata was described as a Lauawania, but an examination of
arr. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—MALLOCH 21
the type sent to me by Doctor de Meijere discloses that it belongs here. I do not know datifrons Thomson which Frey lists in this genus. He suggests its being the same as ocellaris Kertesz, but does not record it from the Philippines. As already indicated I have not seen cyaneos- toma Frey, which is a black species with a large yellow pleural mark and pale bases to the tarsi, and suspect that it does not belong here. It is from the Philippines.
Genus MELANOPACHYCERINA Mailoch
This genus is distinguished from Pachycerina by the possession of 4 pairs of dorsocentral bristles (1+), and a preapical anteroventral comb on the fore tibia. There are three species assigned to the genus as below.
KEY TO THD SPECIES
1. Ocellar bristles quite large, divergent and forwardly directed; frontal orbits glossy black, sharply differentiated from the velvety black interfrontalia, narrowest just in front of ocelli, and gradually widened to anterior margin at which point each is wider than the space between them; fore femur with at least two bristles on apical half of posteroventral surface.
leucochaeta (de Meijere). Ocellar bristles indistinguishable; interfrontalia not narrower at anterior margin than in front of ocelli; fore femur with one posteroventral bristle_ 2.
2. Face bicolored, black and fulvous, with a conspicuous transverse depression on lower half; posthumeral bristle about as long as the dorsocentrals; fore legs entirely black except the coxae and knees____ albiseta (Hendel).
Face entirely glossy black, evenly and prominently convex; posthumeral bristle very short and weak; fore legs with apical halves of tibiae, and the ecoxae, white, the femora yellow at bases, darkened at apices, basal halves OtsipiacvanGrallcohutately Diack 2. a a eee varipes Malloch.
MELANOPACHYCERINA LEUCOCHAETA (de Meijere)
Pachycerina leucochaeta DE MEIJERE, Tijdschr. v. Wnt., vol. 57, p. 256, 1914.
This species occurs in the Nederland Indies and Formosa, and Doctor Frey records it from the Philippines.
It is a conspicuous black species about 5 mm. in length, with hyaline wings.
The other species occur in the Nederland Indies and should be found in the Philippines. I consider Camptoprosopella angustilim- bata de Meijere is albiseta.
Genus LAUXANIELLA Malloch
One species from Formosa has been assigned here, tenuicornis Malloch. It is very similar in general appearance to M. leucochaeta.
22 PROCEEDINGS OF THE NATIONAL MUSEUM Vou. T4 Genus CALLICLYPEUS Frey
This genus: was distinguished from Pachycerina by Frey princi- pally by the shorter third antennal segment, which is not much over twice as long as wide, and were it not for the fact that the descrip- tion further indicates that the anterior pair of dorsocentral bristles is proximad of the suture I would not hesitate to consider it merely a synonym of that genus as Pachycerina sigillata has a similar short third antennal segment. However, without having access to the geno- type, I can not determine the status of the concept and accept it pro- visionally as distinct.
CALLICLYPEUS BOETTCHERI Frey
Calliclypeus boettcheri Frey, Acta Soc. pro Fauna et Flora Fennica, vol. 56, No. 8, p.. 19, 1927.
This species appears to be very similar to Pachycerina sigillata de Meijere in color, the principal distinction lying in the absence of the pair of black marks on the face in sigillata, only the spots between the antennae and eyes being present. The black spots on the seventh abdominal tergite of stgi//ata are not present in boeticheri, though the abdomen is darkened apically.
Locality—Mumungan, Mindanao.
Genus PHOBETICOMYIA Kertresz
This genus was erected for the reception of Lauxania lunifera de Meijere. Frey, in his recent paper on Philippine Sapromyzidae, has included also Lauxvania ornatipennis de Meijere, and boettcher: Frey. The latter is the type species of Poecilomyza a new subgenus of Homoneura erected in the present paper, io ornatipennis belongs to another subgenus of Homoneura.
I have besides lunifera another Philippine species of Phobeti- comyta before me.
The genus is readily distinguished by the slight, but evident, cen- tral bulbosity of the glossy face, backwardly directed anterior orti- tals, and wing venation. The latter is similar to that of Homoneura.
KPY TO THD SPECIES
Wing with a hyaline fascia at extreme apex, and about four hyaline spots in second posterior cell in addition to the one on outer cross vein (fig. 18).
lunifera (de Meijere).
With a hyaline fascia at about its own width from apex, and no hyaline spots in second posterier cell except the one on outer cross vein (fig. 19).
preapicalis, new species.
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—MALLOCH 23
PHOBETICOMYIA LUNIFERA (de Meijere)
Phobeticomyia lunifera pe Metserr, Tijdschr v. Hnt., vol. 53, p. 134, 1910 (Laurania). s Frey records this species from Port Bauga, Mindanao.
PHOBETICOMYIA PREAPICALIS, new species
In addition to the wing markings being distinct this species differs from lunifera in having the face glossy black, with a faint central vertical yellow stripe. In dwnifera the face is glossy black, with the entire lower margin, and a broad V-shaped mark on middle, yellow.
Length, 3.5 mm.
Type, male, and one paratype, both lacking the third antennal seg- ment, Singapore (C. F. Baker). In author’s collection.
Genus TRYPANEOIDES Tonnoir and Malloch
This genus is distinguished from all others except dfelinomyia Kertesz by the presence of two strong bristles on the mesopleura, one on upper hind margin, and the other, which is directed down- ward, close to middle of disk (the latter sometimes duplicated). From Melinomyia it may be distinguished by the presence of two sternopleurals, and the conspicuously marked wings. The costa is the same as in Homonewra, both orbitals are bent backward, the thorax has 1+ 8 strong pairs of dorsocentral and acrostichal bristles, the abdomen has sparse erect setulose hairs, and at apices of all tergites quite conspicuous bristles; there is always at least one con- spicuous bristle a little beyond middle of anteroventral surface of hind femur; the intra-alar bristle is weak or absent; and the fore femur has no anteroventral comb.
The genotype, guttata Tonnoir and Malloch, is found in New Zealand. Below I present a key to the Oriental species known to me. KEY TO THE SPECIES 1. Abdomen glossy black, entirely without markings; three clear spots on
costa of wing between apices of first and second veins; arisia rather LOTS pp VITO Ca eee ae 5s ry Pa SEO OR ee sas Dy, ye es dL ete Se 2. Abdomen brownish or fuscous, with conspicuous gray-dusted spots or mark- ings, and sometimes with brown spots; at least four clear spots on costa DEGWeehaADIEeS On fESG ANG. SeCONd VelINSe es = ee ee 6.
2. Wing with at least two complete clear fasciae in addition to some isolated GlEARISHOLS [AE AR ROL OS oP SEEN Ea PS as Soe NS AS ES See Fk 33 Wing without uninterrupted clear fasciae, sometimes with transverse series ORE CEES SY COS a ee ee ee eee See ee 5.
3. Wing with two clear fasciae, one just proximad of inner cross vein, and sometimes narrowly enclosing it, and the other at about its own width beyond outer cross vein, terminating on costa before apex of second vein,
the clear mark on costa between the fasciae not extending over second
SVG 101 NN eo! es tl cee AE bene woh Din to Bel saree he op a | bicincta (de Meijere). Wing with three complete clear fasciae, the third one extending across the wing from apex of second vein, between the second one and the apical CLOT Pee SO Cea en ee renee BLP eer I ee NN ae See Pe 4,
24 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
4. A clear fasciform mark on costa between the first and second fasciae extend- ing over third vein and faintly to fourth---_________~_ sumatrana Malloch. No clear mark on costa between first and second fasciae.
tricincta Malloch. 5. Wing with but two clear spots in submarginal cell, one close to inner cross
vein and the other on costa between apices on second and third veins. pulchripennis (de Meijere). Wing with three clear spots in submarginal cell, the additional one being
about one-third from apex of cell__--__----_--~- fenestrata (de Meijere).
6. Not more than 10 dark spots and blotches between fifth vein and margin Of Win gisises ik. ys _ ee es ee 8 ae ser seg aa Gs
Not less than 20 dark spots and blotches between fifth vein and margin Of Wi O --6 eR! ae a ee ne ee ee eee 10.
7. Mesopleura with two discal bristles, one above the other______---------- 8. Mesopleura ‘with only ‘one discal: bristle:=- == eee 9.
8. The four clear spots on costa between apices of first and second veins sub- equal in size and quite large; eight clear spots in first posterior cell in- cluding the large one at apex; length 2.5 mm____ tephritina (de Meijere).
More than four clear spots in marginal cell, some very small and others much larger; about 12 clear spots in first submarginal cell including the large one at apex; Jength A mim 2208 2 se ee major, new species.
9. Two clear spots between apices of Second and third veins against costa, the additional one touching tip of third vein and very small; 14 or 15 clear spots in first posterior cell, including the large one at apex.
hyalipuncta Malloch. One clear spot between apices of second and third veins against costa; about eight clear spots in first posterior cell, including the large one at POR Se ee ce ee ae 2 eee Se octomaculata Malloch. 10. Wing narrower than usual, apical hind margin slightly irregular in outline, outer cross vein distinctly less than half as long as ultimate section of fourth vein, the pale markings on wing pale brown except the anterior portions of the spots along costa, and the narrow hind margin, which are Clea ies = 2 oe oa ss 2 Se ee ee ee morio (de Meijere). Wing much broader than usual, conspicuously irregular in outline along apical hind margin, outer cross vein bisinuate, much more than half as long as ultimate section of fourth vein, all the pale wing markings clear, reduced to mere dots or short streaks on most of disk. trypetiformis (de Meijere).
I have seen but one species of this genus from the Philippines, the above key being based upon material supplied by Doctor de Meijere and collected by Mr. E. Jacobson in Java and Sumatra, and by Doctor Toxopeus in Buru. Possibly species of the genus were sent to the late Dr. M. Bezzi by Prof. C. F. Baker, as I know he did send material in this family and also Trypetidae, with which latter family the species may be readily confused.
Doctor Hendel has stated in a recent paper that fenestrata de Meijere belongs to the genus 7'rypetisoma Malloch. The latter has no discal mesopleural bristle and the costa is the same as in Sapro- myza. There is a close resemblance between the wings of Homoneura picta (de Meijere) and Homoneura trypetoptera (Hendel), and cer-
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—MALLOCH 25
tain species of 7rypaneoides, but I have seen both these species and they belong to Homoneura.
Sapromyza perpunctata Lamb, from the Seychelles Islands is ap- parently a species of this genus related to octopunctata Malloch.
TRYPANEOIDES MAJOR, new species
Female.—Black, with dense yellowish gray dust and marked with dark brown. Face with a transverse brown line on upper part, and a biarcuate line of same color near lower margin which connects with a spot on each parafacial at vibrissal angle; antennae brownish yellow, darker below; palpi yellow, fuscous at apices; frons dark at bases of bristles. Thoracic dorsum and pleura with numerous irregular dark brown marks. Abdomen with rather large pale gray dusted spots on hind margins of tergites, a bristle in each spot except the central one which has a bristle on each side of it. Legs testaceous, femora largely brown basally, hind pair dark brown at extreme tips, hind tibia with a narrow dark brown band near base. Wings dark brown, with numerous hyaline spots, four rather large unequal spots and one or two much less distinct streaks in marginal cell, one spot against costa in apex of submarginal cell, preceded by 4 minute dots, then a large oblong spot the inner extremity of which is almost above outer cross vein, and a smaller spot near base of cell; first posterior cell with 12 or 13 clear spots, some very small; 4 or 5 spots in second posterior cell, and about 8 in discal cell, the latter mostly fused in pairs, none of the spots in either cell touching outer cross vein; the apical spot in anal field isolated, the others connected. Knobs of halteres dark brown.
Arista very short haired; orbitals strong. Thorax as in the other species, but the mesopleura with two discal bristles. Each tergite with one series of long erect bristles. Hind femur with about three anteroventral bristles on apical half.
Length, 4 mm.
Type—Cat. No. 41132 U.S.N.M. Baguio, Benquet Province, P. I. (C. F. Baker).
The type bears the number 19391, which indicates that another specimen was sent to some specialist for identification.
Genus MINETTIA Robineau-Desvoidy
There are three segregates of this genus in the material before me. One of these (Minettiella, new subgenus) has but one well- developed sternopleural bristle, the arista pubescent or short haired, frons entirely shining, and the face flat. The other two segregates have two sternopleural bristles, the frons largely or entirely dull, the arista longer haired, and most of the species have the face with
26 PROCEEDINGS OF THE NATIONAL MUSEUM vou. 74
two slight, but distinct, rounded elevations on lower part. The last two segregates I retain in Minettia, the one with the facial eleva- tions being the typical form. In the following key I include all species known to me from the Orient, some of them being dependably separable only by the structure of the male hypopygia.
=~
KEY TO THE SPECIES
. Sternopleura with but one distinct bristle; arista pubescent or very short
haired; frons entirely shining; face flat (Minettiella, new subgenus)-- 2 Sternopleura with two distinct bristles; arista distinctly haired, the shortest hairs at least as long as half the width of third antennal segment; frons largely dull; face usually with two slight elevations below_--------- 3.
. Dorsocentral bristles 1+3, acrostichals 1+4; mesopleura with an isolated an-
terior bristle: below middle 245i.) 2) ._ eee atratula (de Meijere). Dorsocentral bristles 0+2 or 0+3, acrostichals the same; mesopleura with- out an isolated anterior bristle below middle____---~ atrata (de Meijere).
. Face without distinguishable elevations below; male hypopygium with a
stout, tubelike, downwardly projecting dentral process, and without heavily chitinized inner hooks (Fig. 20) ; arista short haired_-------__ 4, Face with a quite noticeable elevation on each side below; male hypopy- gium without a tubelike central process, usually with two or four strong black chitinous hooks or spines, which are more or less curved, and generally asymmetrical: arista, plumose.-2.-—==- =.= 5.
. Large species, fully 5 mm. in length; hind tarsus with the basal segment
slightly compressed in both sexes; ocellar bristles about twice as long as anterion-orbitalsias). tit) Seb a ee 5 eae ee tubifera Malloch. Smaller species, about 4 mm, in length; hind tarsus with basal segment nor- mal in structure in both sexes; ocellar bristles about as long as anterior COT En Sa ie ee ee ee hoozanensis Malloch.
. Bases of wings quite distinctly blackened; knobs of halteres black; abdomen
with distinet ‘dusting ‘on’ dorsum 20) 2 22 See eee 6. Bases of wings not blackened; knobs of halteres black; abdomen entirely shining black, without distinct dusting; hypopygium as Figures 21 and 22. nigrohalterata Malloch.
Bases of wings not darkened; knobs of halteres yellow; abdomen shining black, with grayish dusting evenly distributed__.____________-_-___-___- 8.
. Abdomen reddish testaceous, with evenly distributed grayish dusting.
rufiventris (Macquart). Abdomen black, with grayish dusting, and a dark brown transverse band in, middle. ofeach) tergites: ee ee ee eee eee ii
. Hind tibia with a distinct preapical dorsal bristle; male hypopygium as
WUTC: Dee a a es ae ee ee ee fuscofasciata (de Meijere). Hind tibia without a preapical dorsal bristle; male hypopygium as Figure 24. quadrispinosa Malloch.
. Wings grayish hyaline; anterior one of the postsutural pairs of dorsocentrals
as close to suture as posterior pair is to hind margin; hypopygium as Mipare) 20a ee ee, obscura (de Meijere). Wings, halteres, and squamae and their fringes, honey yellow; anterior pair of postsutural bristles much farther from suture than posterior pair is from“ hind Mmarpints TW eT ee Se ee ee eae 9.
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—-MALLOCH 27
9. Basal portion of male hypopygium (eighth tergite) with the apical lateral arms hinged and furecate, or with internal tooth (Fig. 26).
luteitarsis (de Meijere).
Basal portion of male hypopygium with the apical lateral arms connected
with the basal part by a narrow neck, not hinged to it, and sharp at apex
OT 27) EE 2 So Se ee ea oe philippinensis, new species.
MINETTIELLA, new subgenus
This subgenus is erected for the reception of two species, both of which are glossy black, with immaculate wings, yellow halteres, and characters as given in key.
Subgenotype—Lauxania atratula de Meijere.
Neither species is amongst those before me from the Philippines, though it is very probable that one or both may occur there.
Subgenus MINETTIA Robineau-Desvoidy
In his paper on Philippine Sapromyzidae Doctor Frey gives the sub- generic name H'uminettia to the segregate of Minettia in which there are no elevations on the face, naming lupuliéna Fabricius as genotype. T rather doubt the possibility of distinguishing the segregates by this ‘character, as the elevations, while quite noticeable in some species, are very inconspicuous in others and it is very difficult, if not impossible, to draw the line anywhere. The same author erects the subgenus Calominettia for the South American species geminata Fabricius. The latter he distinguished from H'wménettia by the divergent apical scutellar bristles, a rather variable character.
There are but two species of those listed in the key amongst those before me from the Philippines, though undoubtedly more occur there.
MINETTIA RUFIVENTRIS (Macquart) Minettia rufiventris MacquarT Dipt. Exot. Suppl. 3, p. 68, 1847.
This species appears to be very widely distributed in the Orient, occurring in Formosa, the Nederland Indies, etc. I have before me one specimen from Imugin, N. Viscaya (C. F, Baker), and Frey records it from Luzon.
MINETTIA PHILIPPINENSIS, new species
Male and female—Similar to luteitarsis (de Meijere) in coloration. Black, the face and frons whitish dusted, the former most densely so ; thorax with brownish dust and four faint vittae on dorsum; abdomen shining, without dusting; wings yellowish hyaline, almost honey yellow; halteres honey yellow. The legs are pitchy black, with the bases of tibiae slightly yellowish and the tarsi yellow.
Structurally similar to duteitarsis, the frontal bristles all strong, arista with long hairs; thorax with three pairs of dorsocentrals, the
28 PROCEEDINGS OF THE NATIONAL MUSEUM VOL. 74
anterior pair well behind the suture, and shorter than the second pair; lower anterior part of mesopleura with the hairs long and setalose. Hypopygium as Figure 27.
Length, 6 mm.
Type, male and allotype, Mount Maquiling, Luzon Province; para- type male, Butuan, Mindanao (C. F. Baker).
The species of this genus in the Orient present very good charac- ters in the structure of the male hypopygia for their separation, but these characters have not been used to any extent in describing the species from this or any other region and it is difficult to distinguish some of them on color alone, as they are very similar in appearance. A comparison of the figures of the hypopygia in this paper indicates, if the structure of these organs indicates anything, that there are four groups in the lot: tubifera, nigrohalterata, and philippinensis, representing separate types, and obscura, quadrispinosa, luteitarsis, and fuscofasciata another. Only tubifera lacks the facial elevations. It is noteworthy that the American species with elevations on the face, and also the genotype of Minettia, have entirely different hypo- pygia from this oriental group, there being no heavily chitinized inner hooks present. It is, of course, impossible to find correlated genital characters in the females, so that no groups can be based upon the structure of the male hypopygia, no matter how distinctive this
may be. Genus SAPROMYZA Fallén
I have before me at this time 16 oriental species of this genus which is not so abundantly represented here as in some other regions. Of these species one belongs to a segregate which I consider is en- titled to subgeneric distinction, and it is treated thus herein. To facilitate identification of the species I present a diagnostic key.
KEY TO THE SPECIES
1. Fore femur with an anteroventral comb of minute black bristles apically ; thorax black, with two broad submedian white-dusted vittae, and three pairs of strong dorsocentral bristles, the anterior pair close to suture; abdomen black, with a yellowish dorsocentral line on apical three or four tergites, and apices of all tergites white dusted; wings grayish hyaline, base and outer cross vein slightly fuscous clouded.
albocincta (de Meijere).
Fore femur with an anteroventral comb of minute black bristles apically ;
thorax yellow, with four pairs of dorsocentral bristles (1+3) ; abdomen
yellow, with a large blackish mark on each side of each tergite except the
basal one, which gives the dorsum the appearance of having a broad
blackish vitta on each side; wings yellowish hyaline, outer cross vein and apices of veins 2, 3, and 4, conspicuously clouded with fuscous.
omei, new species.
Fore femur without an anteroventral comb of minute bristles; thorax and
abdomen not colored as above; outer cross vein not clouded__--------- 2.
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—-MALLOCH 29
=I
. Thorax with three pairs of dorsocentral bristles_________-_-__---__----_- 3.
Thorax with two pairs of dorsocentral bristles.-_---.-- +. =+-_-.=.---__ 6.
. Wings broadly brown on costa from base to apex of third vein; species
shining yellow in color, abdomen with a large deep black spot on each side of each tergite; thorax with four fuscous vittae, the submedian pair ex- tending over the sides of scutellum, the others on notopleural sutures, and a fuscous streak over upper margin of sternopleura; legs yellow, with the apices of all femora, all of fore tibiae and tarsi, and extreme
bases of mid and hind tibiae, fuscous______----___-_~ conspicua Malloch. Wings entirely hyaline; species not marked as above_-__-__-________-_-_ 4, . A deep black mark between base of each antenna and eye; ocellar bristles small; prescutellar acrostichals strong______._________ deceptor Malloch. No black mark between each antenna and e¢ye.——-——) 22 5.
. Anterior orbital bristles not much shorter than posterior pair and but little
closer to them than the latter are to vertical bristles, the inner vertical pair not nearly twice as long as outer pair, ocellars minute; a small fuscous spot over ocelli; thorax without a conspicuous dark dorsocentral vitta, the prescutellar acrostichals lacking; abdomen with four black spots on fifth and other four on sixth tergite; wings glassy. hyalipennis (de Meijere). Anterior orbitals minute and much closer to posterior pair than latter are to the verticals, inner verticals about twice as long as outer pair, ocellars long; a large obcordate velvety black mark extending from ocelli back- ward over the occiput, the vertex rounded; thorax with a complete, broad, black, dorsocentral vitta filling area between the dorsocentral series; abdomen largely brown; wings dull hyaline__ koshunensis Malloch.
. Antennae deep black, apex of third segment fulvous yellow, sharply con-
trasting with the basal two-thirds; hairs on arista about as long as width ‘ofsthirdrantennwll seam enti os as Sasa cee eel esl ee inversa, new species. Antennae either yellow or fuscous, the third segment never distinctly paler at apex than at base, usually the reverse, if largely pale at apex the arista
iSsmuUchiGhorbernh wine dates s+ eestor tee + wT ey te ta Nl eh ee 6a.
. Longest hairs on arista never half as long as width of third antennal seg- EN TN a a I a ee ee aS ONS SU ON cl A SE RN EM a Sd Ge Longest hairs on arista at least.as long as width of third antennal seg UTNE) eae ee et LOE AR | OATS APA NS. PE a BE ee A el A A 10.
. Pleura entirely yellow, dorsum of thorax fuscous, with grayish dusting;
hairs on arista black, much denser than usual, the longest about half as
long as the width of third antennal segment; thorax with a pair of dis-
tinct prescutellar acrostichals; antennae with basal segments, and the extreme base of third segment, black, rest of latter yellow.
flavopleura Malloch.
Pleura black or fuscous, concolorous with dorsum of thorax, usually the
hairs on arista not dense and not half as long as width of third antennal
. Thorax with a pair of well aeisona preseutellar acrostichal biletles:
head black, frons in front, and base of third antennal segment broadly, bright orange-yellow; thorax black, with slight brownish dusting on LOTT Ua Ta aa ce a Br nee re PD eB fasciatifrons Kertesz. Thorax without a trace of prescutellar. acrostichal bristles; head and antennae entirely dull yellow; thorax black, with a broad central stripe of yellow dust which extends over disk of scutellum__________________ 9.
5 ADH NERS GATT eS EA ANY 021) Ke al RS tc pollinifrons Malloch
Tibiae darkened at bases and apices_________________- annulifera Malloch.
30 PROCEEDINGS OF THE NATIONAL MUSEUM vou. 74
10. Thorax without distinct prescutellar acrostichal bristles; basal two antennal
segments deep black, third segment yellow__--__-__-___-_____--___- hc
Thorax with a pair of distinct prescutellar acrostichal bristles; basal two
segments of antennae not or but little darker than apical part of third_ 12.
11. Thorax testaceous, dorsum densely whitish gray dusted, with two faint
' dark submedian vittae; apex of scutellum broadly testaceous,
pusillima (de Meijere).
Thorax yellowish gray, gray dusted, with a broad blackish gray vitta be- tween the dorsocentral series which extends on to the scutellum,
pleuralis (Kertesz)
12. Frons fulvous yellow, with a broad dark brown central vitta; antennae deep
black, the base of third segment broadly fulvous yellow; thorax fulvous
yellow, with six broad dark brown vittae on dorsum, the outer one on
each side on the notopleural suture, and the pletra with two similar
Wittaelint sli salu inel fet ie it te SE maquilingensis, new species.
Frons ;and ‘thorax not) colored’ as\abovesii=_ Ss ee ee 13.
18. Mesopleura with a conspicuous downwardly directed bristle near middle of
disk; wing with a black spot at apex of auxiliary vein, and the base
black; thorax black: abdomen testaceous, with six series of dark spots,
which are most distinct on fifth tergite (Xenosapromyza, new subgenus).
cinctipes (de Meijere).
Mesopleura’ with only ‘fine hairs*on disk 21222 hees Sore eee ea 14.
14. Wing with a dark spot at apex of auxiliary vein; basal two antennal seg-
ments and base of third yellowish testaceous, sharply contrasting with
apex of latter which is dark; thorax testaceous, dorsum with a central
vitta and some sublateral marks brown; scutellum testaceous, with two
brown discal marks; abdomen with six series of black spots; femora
fuscous, with preapical yellow annulus_____~_ poecilogaster (de Meijere).
Wings without a dark spot at apex of auxiliary vein; only the base of third
antennal segment yellowish, remainder of antennae fuscous; thorax
fuscous, dorsum densely and uniformly pale gray dusted; scutellum gray
at base, broadly black apically; abdomen with two or four series of black
SpoOtsusil Peres ee ee ee ee ee quadrangulata (de Meijere)-
I have examined all of the Javanese species described by Doctor de Meijere, but only one of them has been received from the Philip- pines. The only other Philippine species known to me are the two described below.
XENOSAPROMYZA, new subgenus
This subgenus is distinguished from Sapromyza by the presence of one or two strong bristles near middle of mesopleura much like those in the genus 7’rypaneoides, but other respects it agrees closely with Sapromyza.
Subgenotype.—Lauxania cinctipes de Meijere.
SAPROMYZA (XENOSAPROMYZA) CINCTIPES (de Meijere)
Lauxania cinctipes DB Mpivrere, Tijdschr v. Ent., vol. 538, p. 125, 1910.
I have seen only the type specimen of this species, from Java. Doctor Frey records the species as a Homoneura from Los Banos, bnt I think erroneously.
ART. 6 NOTES ON ORIENTAL SAPROMYZID FLIES—MALLOCH 31 SAPROMYZA (SAPROMYZA) QUADRANGULATA (de Meijere)
Laurenia quadrangulata pe MrErp, Tijdschr v. Ent., vol. 67, p. 48, 1924.
T have before me a specimen which evidently belongs to this species. It agrees with the type specimen, but is smaller, and has the legs less conspicuously banded, and the abdomen with smaller spots, and with four or even six series instead of but two as in the type. This last character may be variable, as there are traces in the type of additional series of spots besides the two submedian series, and the Philippine example is not fully matured, so accurate comparison is difficult.
Locality, Mount Maquiling, Luzon (C. F. Baker).
SAPROMYZA (SAPROMYZA) MAQUILINGENSIS, new species
Female—Shining fulvous yellow. Frons with a broad, central. dark brown, or fuscous, vitta which covers the ocellar region and extends almost to anterior margin; face with a brownish transverse central mark which does not extend over parafacials; basal two an- tennal segments and apical third of third segment deep black; palpi black. Thoracic dorsum with six, pleura with two, dark brown or fuscous vittae; base of scutellum black in center. Abdomen dark- ened above centrally. Legs yellow, apices of fore femora on anterior side, and all of fore tibia and tarsus, fuscous. Wings yellowish hyaline. Halteres yellow.
Anterior pair of orbital bristles shorter than posterior pair, not as long as the slender ocellars; arista short plumose. Thorax with two pairs of postsutural dorsocentrals, one pair of prescutellar acrostichals, and six series of intradorsocentral hairs, the latter con- fined to anterior portion; no strong bristles on anterior part of mesopleura; both sternopleurals present. Fore femur without an anteroventral comb; mid tibia with one long apical ventral bristle. Inner cross vein close to middle of discal cell; ultimate section of fourth vein fully 2.5 as long as penultimate section.
Length, 3 mm.
Type.—Cat. No. 41161, U.'S.N.M. Mount Maquiling, Luzon (C. F. Baker).
SAPROMYZA (SAPROMYZA) INVERSA, new species
Female—Shining testaceous yellow. Frons broadly black across middle, and on the ocellar spot; antennae deep black, apical third of third segment bright fulvous yellow; aristae black at bases, paler beyond; palpi black, yellow at bases. Thoracic dorsum broadly in- fuscated, the lateral and posterior margins yellow; scutellum with